lunes, 26 de febrero de 2024

V4.2 A Key to the Eleodes Species of the United States and Canada

Some minor changes with Pseudeleodes, E. arcuata, E. debilis, and a few other species. More info in my profile: https://www.inaturalist.org/people/6449260

The key on Google Drive: https://drive.google.com/file/d/1uuMX-OOgYW6vbga-W4rWX2lGJkDtDfmI/view?usp=drivesdk

Publicado el lunes, 26 de febrero de 2024 a las 10:23 AM por eleodesthermopolis eleodesthermopolis | 0 comentarios | Deja un comentario

viernes, 16 de febrero de 2024

V4.1 A Key to the Eleodes Species of the United States and Canada OUTDATED

I’ve made some small improvements both with the keys and the definitions. More info in my profile: https://www.inaturalist.org/people/6449260

The key on Google Drive: https://drive.google.com/file/d/1VsOjMe6kN9xAlW_E7diVmgYn8yuokojq/view?usp=drivesdk

Publicado el viernes, 16 de febrero de 2024 a las 11:34 PM por eleodesthermopolis eleodesthermopolis | 0 comentarios | Deja un comentario

A Key to the Amphidorini Species of Kansas, Nebraska, South Dakota, North Dakota, Missouri, Iowa, Wisconsin, Saskatchewan, and Manitoba

I have created this key to aid in identifying Amphidorini from the northeasternmost part of their range. Kansas, Nebraska, South Dakota, North Dakota, Missouri, Iowa, Wisconsin, Saskatchewan, and Manitoba together have only a total of eleven species, Wisconsin is known only to have Eleodes tricostata. This is likely the easiest key I have created and will create (at least in relation to Amphidorini), and while that is the reason I made it, I will continue to make more regional keys.

The key on Google Drive: https://drive.google.com/file/d/1xWVrT-59QtDlxS_-98GXRKcrws18XWgp/view?usp=drivesdk

1 Elytra carinate…2
1’ Elytra not carinate...6
2 (1) Elytral carina strongly elevated to form a dish...Embaphion muricatum
2’ Elytral carina not strongly elevated...3
3 (2’) Elytra covered in minute setae, luster dull, size less than 23mm…4
3’ Elytra glabrous and with red line running down back, luster usually shiny, size more than 23mm...5
4 (3) Body oval shaped, elytra without rows of tubercles...Eleodes opaca
4’ Body not oval shaped, elytra with rows of tubercules…Eleodes tricostata
5 (3’) Pronotum concave...Eleodes suturalis
5’ Pronotum convex…Eleodes acuta
6 (1’) Size 18mm or larger...7
6’ Size smaller than 18mm…9
7 (6) Body elongate, elytra smooth, never sulcate, profemora never dentate…Eleodes longicollis
7’ Elytra sulcate, profemora dentate in males…8
8 (7’) Anterolateral angles of pronotum acute…Eleodes hispilabris
8’ Anterolateral angles of pronotum rounded…Eleodes obscura obscura
9 (6’) Elytral punctures in rows and less numerous, elytra often with ridges or a red line…Eleodes carbonaria obsoleta
9’ Elytral punctures numerous, elytra never with ridges or a red line…10
10 (9’) Hind angles of pronotum wider than anterolateral angles...Eleodes fusiformis
10’ Hind angles and anterolateral angles of pronotum about the same width…Eleodes extricata

Publicado el viernes, 16 de febrero de 2024 a las 10:54 PM por eleodesthermopolis eleodesthermopolis | 0 comentarios | Deja un comentario

A Key to the Amphidorini species of Southern California

In this journal post (https://www.inaturalist.org/journal/eleodesthermopolis/87122-a-key-to-the-eleodes-species-of-southern-california-outdated) I created a key for the Eleodes of southern California, I had a more limited definition of the location at that time, and this key aims to not only improve what I had done in that key, but also expand it to all of Amphidorini, and the more common definition of southern California. I have been told by M. A. Johnston that regional keys are good, which I certainly would agree (while technically a regional key, my Eleodes key for the U.S. and Canada has more or less 80 outcomes, the help in regional keys is primarily the lower species count, and potentially less variation), anyway, I do plan to make more regional keys across the range of Amphidorini, so far, there is only this one, and another which I will soon post for many states/provinces in the northeastern part of the tribe’s range.

The key on Google Drive: https://drive.google.com/file/d/1ZqjZsoLCLD6hM5coFTku25erj4e7HUTd/view?usp=drivesdk

1 Elytra with at least four large costae, size small, pronotum with strongly projected anterolateral angles, tuberculate or reticulate, arid regions…2 (genus Trogloderus)
1’ Elytra lacking costae...5
2 (1) Pronotum tuberculate…3
2’ Pronotum reticulate…4
3 (2) Both elytra with a large tubercule at the apex, pronotum broader…Trogloderus tuberculatus
3’ Elytral apex without two large tubercules, pronotum less broad…Trogloderus skillmani
4 (2’) Pronotum less broad and clearly and evenly convex…Trogloderus major
4’ Pronotum more broad and while still convex, not as even…Trogloderus vandykei
5 (1’) Elytra carinate, size small, arid regions…Embaphion depressum
5’ Elytra never carinate…6
6 (5’) Entire body covered in long conspicuous setae, size less than 15mm…7
6’ Never with conspicuous setae unless size is over 15mm, or said conspicuous setae long, golden, and primarily from beneath the sides of the pronotum, or the pronotum is strongly cordate…11
7 (6) Size 13mm or more...8
7’ Size 12mm or less…9
8 (7) Setae usually orange, pronotum constricted at base…Eleodes osculans
8’ Setae black or grey, pronotum not constricted at base, Baja California…Eleodes ursus
9 (7’’) Setae black, pronotum constricted at base…Eleodes nigropilosa
9’ Setae yellow, pronotum not constricted at base…10
10 (9’) Antennomere 3 subequal to 4, more northern from the U.S.-Mexico border…Eleodes littoralis
10’ Antennomere 3 subequal to 4 and 5 combined, more southern from the U.S.-Mexico border…Eleodes subdeplanata
11 (6’) Elytra with many clearly separated tubercules, apex attenuate, pronotum legs long when compared to the body, usually near mountains…Eleodes granosa
11’ If elytra tuberculate, then tubercules usually not as clearly separate, and legs never abnormally long when compared to the body…12
12 (11’) Long, golden setae originating from beneath the sides of the pronotum, Mohave Desert…genus Lariversius
12’ If with conspicuous setae, then setae brownish…13
13 (12’) Body appearing rough, elytral punctures conspicuous and never simple, pronotum broad, only slightly constricted at base, elytra strongly attenuate, San Luis Obispo and Santa Barbara counties…Eleodes marginata
13’ If pronotum broad then elytral punctures simple and minute...14
14 (13’) Elytra sulcate…15
14’ Elytra not sulcate…16
15 (14) Anterolateral angles of pronotum acute…Eleodes hispilabris
15’ Anterolateral angles of pronotum rounded…Eleodes obscura sulcipennis
16 (14’) Elytra with clearly separated papillae, pronotum arcuate, about the same width as the elytra, foretibia curved, buccal process strongly projected, in and around San Diego county…Eleodimorpha bolcan
16’ Buccal process not strongly projected…17
17 (16’) Mesofemora and metafemora dentate or subdentate...18
17’ Mesofemora and metafemora never dentate…20
18 (17) Pronotum widest anterior of middle, elytra with simple punctures, widespread…Eleodes armata
18’ Pronotum widest at middle, elytral punctures simple or muricate, Baja California…19
19 (18’) Elytra usually appearing rough, metatibia more or less straight…Eleodes moesta
19’ Elytra appearing smooth, metatibia curved or bent…Eleodes femorata
20 (17’) Pronotum evenly arcuate, widest at middle, elytra with fairly conspicuous brownish setae, muricate punctures, luster dull, larger than 18mm, Channel Islands…Eleodes adumbrata
20’ Elytra never with conspicuous setae unless the pronotum is strongly cordate or size is less than 18mm…21
21 (20’) Profemora always dentate, size larger...22
21’ Profemora never dentate, size variable...28
22 Pronotum broad, elytral apex never attenuate, rounded, humeri obsolete...Eleodes grandicollis
22’ If pronotum broad, then elytral apex attenuate or caudate, humeral angles never obsolete...23
23 (22’) Pronotum at least two times the width of the head, profemora always dentate, often caudate...Eleodes acuticauda
23’ Pronotum less then two times the width of the head...24
24 (23’) Pronotum widest anterior of middle...25
24’ Pronotum widest at middle...26
25 (24) Pronotum usually broad and strongly arcuate, striae more dense but punctures smaller, usually in more northern forests…Eleodes dentipes
25’ Pronotum more parallel sided, only slightly arcuate, striae spaced out but punctures larger...Eleodes gracilis distans
26 (24’) All tibias strongly curved, profemoral spines blunt, never caudate...Eleodes subcylindrica
26’ Tibias not abnormally curved, profemoral spines acute, caudate or not...27
27 (26’) Elytral punctures minute, appearing inpunctate, inland chaparral...Eleodes discincta
27’ Elytral punctures small but fairly conspicuous, arid regions...Eleodes mexicana
28 (21’) Punctures of elytra simple, elytra never with tubercules or papillae, pronotal punctures always small and minute...29
28’ Punctures of elytra not simple, elytra with or without tubercules or papillae, pronotal punctures variable...33
29 (28) Body elongate, size over 22mm...Eleodes gigantea
29’ Body elongate or not, size 22mm or less...30
30 (29’) Body not particularly elongate, legs more robust...31
30' Body elongate, legs more slender, often long, aid regions...32 (Eleodes (Metablapylis))
31 (30) Strial punctures larger than those of intervals, Mohave Desert near Nevada...Eleodes carbonaria carbonaria
31’ Strial punctures and those of intervals subequal, chaparral and adjacent desert...Eleodes carbonaria omissa
32 (30') Strial punctures larger than those of intervals, pronotum more arcuate...Eleodes californica
32’ Strial punctures and those of intervals subequal…Eleodes delicata (Eleodes nevadensis)
33 (28') Pronotum usually strongly cordate, elytral apex usally rounded, the coast and forests...Eleodes (Blapylis)
33' Pronotum evenly arcuate, elytral apex usually more or less attenuate, mountain forests...Eleodes (Litheleodes)

Publicado el viernes, 16 de febrero de 2024 a las 10:44 PM por eleodesthermopolis eleodesthermopolis | 0 comentarios | Deja un comentario

miércoles, 14 de febrero de 2024

V4.0 A Key to the Eleodes Species of the United States and Canada OUTDATED

While I have made a few minor changes in the keys, this new version, now V4.0, is primarily a change in formatting and includes more help in identification. I have changed it to two columns on each page, and have added the number of the previous couplet on the one you’re currently on. I have also added some definitions to certain words (let me know if I should alter them), and I’ve added a few figures. Since this update includes more than just keys, I’ve decided that the Google Drive link is now the only way that this version will be accessed (instead of also on the journal post itself). More info in my profile: https://www.inaturalist.org/people/6449260

The key on Google Drive: https://drive.google.com/file/d/1SLOCrU6UJ73To81TOY5N9mOd0SnIROZG/view?usp=drivesdk

Publicado el miércoles, 14 de febrero de 2024 a las 11:01 AM por eleodesthermopolis eleodesthermopolis | 0 comentarios | Deja un comentario

domingo, 11 de febrero de 2024

V3.6 A Key to the Eleodes Species of the United States and Canada OUTDATED

Some fairly major changes have been made. The Promus section (37 to 41) has had some adjustments to aid with more diverse forms of its species, and E. nigrina (continuation) and E. inornata have been moved from around couplet 44, to couplets 81 and 82, this should help with distinguishing from E. humeralis, which has also been altered. I have also made a few small miscellaneous changes. More info in my profile: https://www.inaturalist.org/people/6449260

The key on Google Drive (looks nicer and can be saved): https://drive.google.com/file/d/1eFmmPiPeA57nQSG4Lhp-saTloFgfcFMI/view?usp=drivesdk

1 Mesofemora and metafemora dentate or subdentate...2
1’ Mesofemora and metafemora never dentate...4
2 Pronotum widest anterior of middle, elytra with simple punctures, widespread...E. armata
2’ Pronotum widest at middle, elytral punctures simple or muricate, Baja California...3
3 Elytra usually appearing rough, metatibia more or less straight...E. moesta
3’ Elytra appearing smooth, metatibia curved or bent...E. femorata
4 Size small, pronotum generally cordate and constricted at base, elytral apex not attenuate or caudate, roughly sculptured, never sulcate, sometimes with short setae, usually distributed across beaches, forests, and mountains, rarely in arid regions...subgenus Blapylis
4’ Size variable, pronotum usually not cordate, if so then punctures smaller and more minute, pronotum often slightly constricted at base, elytral apex variable, roughly sculptured, sulcate, setose or not, widespread across all environments...5
5 Legs long with an unproportionately small body, robust or not, body often flattened if not robust, never sulcate, if legs shorter, then body elongate or robust and flattened, size smaller...6
5’ Legs long or short but with body fairly proportionate, only rarely flattened, sulcate or not, if legs fairly long, then not with conspicuous setae, or a small, elongate, and flattened body, size variable...22
6 Body generally slender or rarely robust, always flattened, pronotal and elytral sculpturing simple (except in E. nigrina), elytra usually more parallel sided, never with conspicuous setae...7
6’ Body, generally robust, never flattened, pronotal and elytral sculpturing complex, elytra usually more ventricose, usually with conspicuous setae...18
7 Elytral sculpturing muricate or scabrous, southern New Mexico to Canada, Sierra Nevada Mountains to Rocky Mountains...E. nigrina (in part)
7’ Elytral sculpturing simple, always below the northernmost edge of Colorado’s border...8
8 Range only in the Inyo White Mountains...9
8’ Range widespread but not in the Inyo White Mountains...10
9 Profemora with blunt tooth, anterolateral angles of pronotum blunt, but projected, legs shorter...E. aalbui
9’ Profemora only sinuate, anterolateral angles of pronotum slightly projected, legs longer...E. microps
10 Range near western Texas and the Guadalupe Mountains...11
10’ Range in the Southwest United States excluding Texas and the Guadalupe Mountains...13
11 Apex of the the probasitarsus with a brush of golden setae, range in the Guadalupe Mountains...E. guadalupensis
11’ Probasitarsus lacking golden setae, only with orange spines, range near southwestern Texas...12
12 Mentum with finger-like process at the center of its frontal margin, pronotum lacking projected anterolateral angles, widest at middle, near Rio Grande river...E. labialis
12’ Mentum without finger-like process, pronotum with projected anterolateral angles, Big Bend National Park...E. easterlai
13 Range in the Mohave and Colorado Deserts of California, pronotum usually evenly arcuate, strial punctures larger than those of intervals...E. californica
13’ Range in Arizona, Utah, and the Colorado Desert, pronotum rarely evenly arcuate, strial punctures variable, but if in the Colorado Desert, then punctures of striae and intervals small and subequal in size...14
14 Anterolateral angles of pronotum strongly projected, caves near the Arizona-Utah border...E. wynnei
14’ Anterolateral angles of pronotum not projected or only slightly projected, if strongly projected then not near the Arizona-Utah border...15
15 Strial punctures medium sized, larger than those of intervals…16
15’ Strial punctures smaller, subequal to those of intervals…17
16 Anterolateral angles of pronotum projected, Sky Islands in southeast Arizona...E. dissimilis
16’ Anterolateral angles not projected, truncate, the Grand Canyon...E. leptoscelis
17 Size smaller, Sonoran Desert including the Colorado Desert...E. delicata (E. nevadensis)
17’ Size larger, northern Arizona and southern Utah, usually in forested areas...E. wheeleri
18 Fairly large tubercles present on elytra...19
18’ Elytra without large tubercules, or tubercules quite small...20
19 Range in southern California to Oregon...E. granosa
19’ Range near the Four Corners...E. leechi
20 Pronotum evenly tuberculate, Oregon...E. spoliata
20’ Pronotum punctate, widespread...21
21 Antennas robust, body generally more robust, setae shorter...E. hirsuta
21’ Antennas thin, body generally more slender, setae longer...E. pilosa
22 Size small or sometimes medium, covered in long setae, restricted to coastal and sometimes inland California, or rarely in western Nevada, or the Sonoran Desert...23
22’ Size small or large, usually glabrous, if with conspicuous setae then not found in coastal California, nor in Baja California, or the Sonoran Desert, and if found in western Nevada then third antennomere not shortened...28
23 Males with dentate profemora and elytra caudate, Sonoran Desert...E. tribulus
23’ Males nor females ever caudate or attenuate, non-arid California and/or western Nevada...24
24 Third antennomere shortened, body smaller and more slender, pill shaped, setae yellow, coastal California and towards and into western Nevada...E. littoralis
24’ Third antennomere not shorted, body larger, California...25
25 Size 13mm or more...26
25’ Size 12mm or less...27
26 Setae usually orange, pronotum constricted at base, California and Baja California...E. osculans
26’ Setae black or grey, pronotum not constricted at base, Baja California...E. ursus
27 Setae black, pronotum constricted at base, California and Baja California...E. nigropilosa
27’ Setae yellow, pronotum not constricted at base, Baja California...E. subdeplanata
28 Elytra caudate at least in males and very rough, near Tamaulipas if not caudate, size usually fairly large...29
28’ Elytra caudate or not, but if caudate and roughly textured then elytra only slightly rough, elytra can be rough if not caudate and not near Tamaulipas, size large or small...31
29 Only males caudate with profemora dentate, Tamaulipas...E. fiski
29’ Both males and females caudate with mutic profemora, Four Corners and Sierra Nevada region...30
30 Elytra lacking long setae, Four Corners...E. caudifera
30’ Elytra always with long setae, Sierra Nevada region...E. longipilosa
31 Elytral sculpture very rough, elytra always attenuate but never caudate, pronotum wide, California or slightly into Nevada or Oregon...32
31’ Elytral sculpture variable but if rough then elytra never attenuate and pronotum never wide, widespread...33
32 Luster shiny and elytral apex acutely attenuate, Pacific coast...E. marginata
32’ Luster dull and elytral apex blunt but still attenuate, Sierra Nevada Mountains...E. scabricula (E. acutangula)
33 Body small and fusiform, punctures usually small and scattered, sometimes with minute setae, pronotum never sinuate before its base...34
33’ Shape rarely fusiform, if so, then punctures larger and in rows, pronotum sinuate before base, or size large...37
34 Elytra carinate and with minute setae…35
34’ Elytra without carina or minute setae...36
35 Line where pronotum and elytra meet nearly a perfect straight line, coastal Texas and Louisiana...E. veterator
35’ Line where pronotum and elytra meet rounded, making the elytra’s humeral angles appear projected, northern Texas to Canada...E. opaca
36 Body more fusiform, hind angles of pronotum quite a bit wider than anterolateral angles...E. fusiformis
36’ Body less fusiform, hind angles and anterolateral angles about the same width...E. extricata (in part)
37 Size medium, elytra inflated, apex rounded, and legs shorter, or elytra fairly elongate, apex slightly attenuate, and legs longer, elytral striae composed of large, moderately spaced, conspicuous punctures, pronotum fairly parallel sided, luster fairly dull, never caudate, Texas and southeast New Mexico...E. goryi
37’ Elytral striae not composed of large punctures, if so then size smaller and luster shiny...38
38 Size medium, elytra usually flattened, always attenuate, and heavily punctate, although punctures in rows, pronotum arcuate, luster shiny, never sulcate, caudate, or carinate, south and central Texas...E. striolata
38’ Size, elytra, punctures, luster, and pronotum variable, though if in south and central Texas then elytra sulcate, caudate, or carinate, or elytra less heavily punctate, widespread...39
39 Size medium, body subfusiform, often droplet shaped, luster usually dull, pronotum widest at middle, parallel sided or arcuate, elytra usually glabrous although rarely with minute setae, inflated or not, apex often slightly attenuate and with a split between both elytra, profemora dentate in males, never sulcate, caudate, or carinate, Arizona, New Mexico, and Texas…40
39’ Size, luster, pronotum, and elytra variable, body rarely fusiform or subfusiform if not quite elongate and lacking dentate profemora, even more rarely droplet shaped, setose, sulcate, caudate, carinate or not, widespread…42
40 Elytra with minute setae and heavily punctate, eastern Texas…E. spiculifera
40’ Elytra glabrous and punctures inconspicuous, Arizona and New Mexico…41
41 Pronotum more or less parallel sided, anterolateral angles almost always projected outwards, prosternum forming large triangular wedge...E. subnitens
41’ Pronotum evenly arcuate, anterolateral angles only rarely projected outwards, prosternum not forming large triangular wedge...E. madrensis
42 Size almost always larger than 19mm, almost always with legs long, no matter the size, sometimes caudate, profemora often dentate, at least in males...43
42’ Size almost always smaller than 19mm, if larger, then legs almost always short unless body is fusiform, legs are often long if body in normal size range, never caudate, profemora only rarely dentate...70
43 Body elongate, elytra sometimes inflated, pronotum usually not all that distinct from elytra (subfusiform), if the pronotum and elytra are fairly distinct, then elytra smooth, prosternum projected downwards or profemora strongly sinuate, profemora never dentate and never with caudate elytra, punctures simple and minute, never sulcate...44
43’ Body fairly variable, sometimes elongate, often robust, pronotum usually quite distinct from elytra, almost always with dentate profemora, if missing then profemora slightly sinuate, elytra sometimes caudate, sulcate or not...47
44 Elytra with red line running down the length of the elytra, this line never diffuses into black...E. ornatipennis
44’ Elytra entirely black, or if a red line is present along the elytra it diffuses into black...45
45 Profemora strongly sinuate, pronotum widest slightly anterior of middle...E. hepburni
45’ Profemora simple, pronotum widest at middle...46
46 Prosternum smaller, only slightly projecting downwards, restricted to California...E. gigantea
46’ Prosternum larger prominently projecting downwards, widespread but not found in California...E. longicollis
47 Elytra with carina, if carina absent and the elytra either sulcate or with numerous irregular punctures, then anterolateral angles of the pronotum rounded off and the pronotum widest anterior of middle, pronotum concave or convex...48
47’ Elytra lacking carina, if elytra sulcate or with numerous irregular punctures, then anterolateral angles of the pronotum acute, or if angles not acute then pronotum widest at middle, pronotum convex...53
48 Pronotum concave...E. suturalis
48’ Pronotum convex...49
49 Elytra with carina...E. acuta
49’ Elytra without carina...50 (E. obscura)
50 Elytra with clear, prominent striae...51
50’ Elytra without clear, prominent striae...52
51 Elytra deeply sulcate, west of the Great Plains...E. obscura sulcipennis
51’ Elytra slightly sulcate, Great Plains...E. obscura obscura
52 Elytra with simple punctures, western Texas and southern New Mexico...E. obscura glabrisulca
52’ Elytra with muricate punctures, Four Corners...E. obscura dispersa
53 Elytra with large glabrous costae, intervals minutely hirsute and often coated in white debris, caudate or not, legs long...E. mirabilis
53’ Elytra variable but without costae, caudate or not...54
54 Pronotum usually wide and strongly arcuate, humeral angles of elytra obsolete, apex rounded...E. grandicollis
54’ Pronotum variable, if wide, then elytral apex attenuate at least, humeral angles of elytra not obsolete...55
55 Elytra strongly sulcate, at least on elytral apex…56
55’ Elytra not strongly sulcate…57
56 Pronotum widest anterior to middle, anterolateral angles of pronotum acute, widespread…E. hispilabris
56’ Pronotum widest at middle, anterolateral angles of pronotum rounded off, Texas…E. composita
57 Restricted to California and Baja California, if not in California then tibia strongly curved, profemoral spine small and blunt, and in Nevada, northwest Arizona, or extreme southwest Utah...58
57’ Widespread, but not found in California, Baja California, Nevada, northwest Arizona, or southwest Utah...64
58 Pronotum two or more times wider than the head, often caudate...E. acuticauda
58’ Pronotum less then two times wider than the head, rarely caudate...59
59 Pronotum more or less parallel sided, widest at middle, Mohave and Colorado Deserts, Nevada and Arizona...60
59’ Pronotum not parallel sided, or if so then elytral apex strongly attenuate and with groove in between both elytra, if in the Mohave or Colorado Desert then pronotum widest anterior of middle…61
60 Tibia strongly curved, profemoral spines blunt, never caudate...E. subcylindrica
60’ Tibia not strongly curved, profemoral spines acute, males caudate...E. mexicana
61 Pronotum widest at middle...62
61’ Pronotum widest anterior of middle...63
62 Elytra glabrous, punctures simple, luster shiny, elytral apex with groove in between the elytra, Baja California…E. discincta
62’ Elytra setose, punctures muricate, luster dull, elytral apex without groove in between the elytra, Channel Islands…E. adumbrata
63 Pronotum usually broad and strongly arcuate, widespread across California but absent near the U.S.-Mexico border…E. dentipes
63’ Pronotum more parallel sided, only slightly arcuate, southern California…E. gracilis distans
64 Body usually flattened, elytral punctures simple, in rows, and more minute, never caudate, pronotum widest at middle, luster dull, Texas…E. knullorum
64’ Body usually not flattened, punctures variable, caudate or not, pronotum variable, if widest at middle then luster shiny, widespread…65
65 Elytral punctures irregular and strongly muricate, never caudate…E. sponsa
65’ Elytral punctures in rows and with simple punctures, or slightly asperate punctures, mainly at the elytral apex, elytra usually caudate, at least in males…66
66 Anterolateral angles of pronotum facing straight forward or only slightly projected outwards, hind angles not rounded, males caudate and with the metatibia sinuate, southern Arizona…E. eschscholtzii eschscholtzii
66’ If in southern Arizona then anterolateral angles of pronotum projecting outwards, or hind angles of pronotum appearing quite rounded…67
67 Hind angles of pronotum appearing rounded, size large, elytra often inflated and strongly convex, or fairly parallel sided…68 (E. spinipes)
67’ Hind angles of pronotum not rounded or only slightly rounded, size medium to large, elytra never inflated, often parallel sided…69
68 Elytra more or less parallel sided, west of the Pecos River…E. spinipes macrura
68’ Elytra ventricose, east of the Pecos River…E. spinipes ventricosa
69 Elytra caudate or strongly attenuate, punctures smaller, El Paso region…E. tenuipes
69’ Elytra at most slightly attenuate, punctures larger…E. gracilis gracilis
70 Elytra with conspicuous setae…71
70’ Elytra glabrous, or only with short inconspicuous setae…75
71 Size less than 12mm, elytra shiny with complex sculpturing and covered in golden hairs, areas with sandy soils around the Four Corners…E. barbata
71’ Size usually small but rarely less than 12mm, elytra usually dull, sometimes shiny, never with golden hairs, usually in deserts above the Sonoran Desert…72
72 Size larger, body more robust, northern Arizona and southern Utah…E. halli
72’ Size smaller, body more slender, generally around the Great Basin or the Great Sand Dunes of Colorado…73
73 Endemic to the Great Sand Dunes National Park…E. hirtipennis
73’ Distributed generally around the Great Basin…74
74 Setae black, generally around the Great Basin…E. letcheri
74’ Setae brownish, in and around Utah…E. aspera
75 Elytra flat and widened, about the same width as and meeting with pronotum to form a continuous line…76
75’ Elytra usually not flat or widened, and never meeting with pronotum to form a line, elytra elongate or ovate…80
76 Body covered in minute setae, elytra with rows of tubercules, widespread…E. tricostata
76’ Body glabrous, elytra lacking rows of tubercules, Texas, Oklahoma, and New Mexico…77
77 Elytral sculpturing rough, striae vague to absent, luster generally dull…E. neomexicana
77’ Elytra with clear striae, luster shiny…78
78 Elytra with deeply impressed striae…E. pedinoides
78’ Elytra with slightly impressed striae…79 (E. wenzeli)
79 Punctures of elytra fine but easily noticeable…E. wenzeli speculicollis
79’ Punctures of elytra extremely fine and minute…E. wenzeli wenzeli
80 Both elytra with around three large smooth costae, legs often dark red, southern Utah…E. parowana
80’ Both elytra without at least three large costae…81
81 Body elongate, elytra appearing smooth with minute muricate punctures, each puncture with a tiny seta arising from it, entire tarsus covered heavily with long setae, anterolateral angles of pronotum projected and acute, western Nevada…E. inornata
81’ Tarsi never entirely and heavily setose, at most only with short sparse setae, or some interrupting the plantar groove, elytra and pronotum variable, but if in Nevada then anterolateral angles not projected and acute…82
82 Elytral punctures muricate or scabrous, usually appearing rough, body elongate, elytra parallel sided and with not particularly attenuate, luster usually dull…E. nigrina (continuation)
82’ Punctures variable, if elytra appear rough then body not particularly elongate, and elytra more arcuate and attenuate…83
83 Size small, elytral luster generally shiny, elytra heavily punctate, usually irregular, never with tubercules, pronotal punctures small and sparse…E. extricata (continuation)
83’ Size small to medium, luster variable although generally dull if elytra with irregular punctures, said irregular punctures also smaller, elytra with punctures in rows or irregular, or with tubercules, papillae, or rugae…84
84 Larger protibial spur extending almost to third tarsomere, pronotum broader than long, elytra with small, granulate tubercules, luster dull, Pacific Northwest except southeast Oregon…E. humeralis
84’ Larger protibial spur never reaching this size if in the region stated above, but may be of this size if outside that region including southeast Oregon, if elytra tuberculate then tubercules larger and luster shiny…85
85 Elytra with papillae, pronotum wide, luster dull, California…E. papillosa
85’ Elytra without papillae, luster variable…86
86 Elytra with coarse, transverse, tuberculate rugae, luster shiny…E. corvina
86’ Elytra without transverse rugae, tuberculate or not, luster variable…87
87 Elytra tuberculate, tubercules larger, elytra noticeably rough, protibial spurs smaller…88
87’ Elytra not tuberculate, or if so then tubercules smaller, elytra less rough, and protibial spurs larger…89
88 Pronotum more or less arcuate and wider than long…E. granulata
88’ Pronotum more or less parallel sided and longer than wide…E. subtuberculata
89 Size small, pronotum evenly arcuate, punctures minute and in rows, luster usually dull, body never elongate, Arizona and New Mexico…E. arcuata
89’ Size variable, pronotum variable, but if evenly arcuate, then punctures more conspicuous, luster variable…90
90 Luster shiny, pronotum evenly arcuate, elytral punctures usually fairly large and in rows, outer protibial spur clearly larger than inner protibial spur, body usually ovate…E. debilis
90’ Luster variable, pronotum variable, punctures variable but rarely large, if protibial spurs disproportionate in size then luster more dull and elytra with tubercules or smaller punctures and a more elongate form…91
91 Longer protibial spur extending to almost to the third tarsomere, elytra with small to moderately sized tubercules, either in rows or irregular…92 (E. rileyi)
91’ Longer protibial spur extending only to apex of the first tarsomere, elytra never tuberculate…93
92 Elytra shiny with prominent striae, Four Corners…E. rileyi reducta
92’ Elytra dull without very prominent striae, widespread…E. rileyi rileyi
93 “…longer protibial spur parallel sided in basal half then tapering to a point at apex,” luster dull, punctures irregular if in California, if not then punctures in rows…94
93’ “…longer protibial spur evenly tapering to a point from base to apex,” luster usually shiny, sometimes dull, punctures variable, if in California then luster usually dull and punctures with some semblance of rows…96 (E. carbonaria)
94 Elytral punctures irregular, San Francisco region…E. quadricollis
94’ Elytral punctures organized in rows, generally around southeast Arizona…95 (E. anthracina)
95 Size larger, elytral punctures less fine, widespread…E. anthracina anthracina
95’ Size smaller, elytral punctures more fine, Chiricahua Mountains…E. anthracina lustrans
96 Strial punctures and those of intervals subequal in size, often diffusing, restricted to California…E. carbonaria omissa
96’ Punctures variable, but if in California then strial punctures larger than those of intervals and in rows…97
97 Body usually fairly fusiform, punctures conspicuous and in rows, east of the Pecos River and always below the southernmost edge of Arizona and New Mexico’s border…E. carbonaria soror
97’ Body only rarely fusiform, punctures variable, west of the Pecos River, or if not then above the southernmost edge of Arizona and New Mexico’s border…98
98 Punctures of elytra simple…100
98’ Punctures of elytra muricate…99
99 Strial punctures and those of intervals subequal in size, often diffusing, generally around northwest New Mexico…E. carbonaria knausii
99’ Strial punctures larger than those of intervals, striae in rows, often in ridges, widespread…E. carbonaria obsoleta
100 Body usually more fusiform, punctures more minute, size larger than 17mm, southern Arizona and New Mexico…E. carbonaria chihuahuaensis
100’ Body usually not fusiform, punctures more conspicuous, size 17mm or smaller, widespread…E. carbonaria carbonaria

Publicado el domingo, 11 de febrero de 2024 a las 03:16 AM por eleodesthermopolis eleodesthermopolis | 0 comentarios | Deja un comentario

jueves, 08 de febrero de 2024

V3.5 A Key to the Eleodes Species of the United States and Canada OUTDATED

Several new changes. I have changed some things around in the Eleodes and Metablapylis section, as well as improved many miscellaneous couplets. More info in my profile: https://www.inaturalist.org/people/6449260

The key on Google Drive (looks nicer and can be saved): https://drive.google.com/file/d/1XJXzO7FNb4bkl60Wg4A1te2v3SU1sEsZ/view?usp=drivesdk

1 Mesofemora and metafemora dentate or subdentate...2
1’ Mesofemora and metafemora never dentate...4
2 Pronotum widest anterior of middle, elytra with simple punctures, widespread...E. armata
2’ Pronotum widest at middle, elytral punctures simple or muricate, Baja California...3
3 Elytra usually appearing rough, metatibia more or less straight...E. moesta
3’ Elytra appearing smooth, metatibia curved or bent...E. femorata
4 Body small, pronotum generally cordate and constricted at base, elytral apex not attenuate or caudate, roughly sculptured, never sulcate, sometimes with short setae, usually distributed across beaches, forests, and mountains, rarely in arid regions...subgenus Blapylis
4’ Body small or large, pronotum usually not cordate, if so then punctures smaller, more minute, often slightly constricted at base, elytral apex attenuate, caudate, or neither, roughly sculptured or not, sulcate or not, with setae or not, widespread across all environments...5
5 Legs long with unproportionately small body, robust or not, body often flattened if not robust, never sulcate, if legs shorter, then body elongate or robust and flattened, size smaller...6
5’ Legs long or short but with body fairly proportionate, only rarely flattened, sulcate or not, if legs fairly long, then not with conspicuous setae, or a small, elongate, and flattened body, size variable...22
6 Body generally slender or rarely robust, always flattened, pronotal and elytral sculpturing simple (except in E. nigrina), elytra usually more parallel sided, never with conspicuous setae...7
6’ Body, generally robust, never flattened, pronotal and elytral sculpturing complex, elytra usually more ventricose, usually with conspicuous setae...18
7 Elytral sculpturing muricate or scabrous, southern New Mexico to Canada, Sierra Nevada Mountains to Rocky Mountains...E. nigrina (in part)
7’ Elytral sculpturing simple, always below the northernmost edge of Colorado’s border...8
8 Range only in the Inyo White Mountains...9
8’ Range widespread but not in the Inyo White Mountains...10
9 Profemora with blunt tooth, anterolateral angles of pronotum blunt but projected, legs shorter...E. aalbui
9’ Profemora only sinuate, anterolateral angles of pronotum slightly projected, legs longer...E. microps
10 Range near western Texas and the Guadalupe Mountains...11
10’ Range in the Southwest United States excluding Texas and the Guadalupe Mountains...13
11 Apex of the the probasitarsus with a brush of golden setae, range in the Guadalupe Mountains...E. guadalupensis
11’ Probasitarsus lacking golden setae, only with orange spines, range near southwestern Texas...12
12 Mentum with finger-like process at the center of its frontal margin, pronotum lacking projected anterolateral angles, widest at middle, near Rio Grande river...E. labialis
12’ Mentum without finger-like process, pronotum with projected anterolateral angles, Big Bend National Park...E. easterlai
13 Range in the Mohave and Colorado Deserts of California, pronotum usually evenly arcuate, strial punctures larger than those of intervals...E. californica
13’ Range in Arizona, Utah, and the Colorado Desert, pronotum rarely evenly arcuate, strial punctures variable, but if in the Colorado Desert, then punctures of striae and intervals small and subequal in size...14
14 Anterolateral angles of pronotum strongly projected, caves near the Arizona-Utah border...E. wynnei
14’ Anterolateral angles of pronotum not projected or only slightly projected, if strongly projected then not near the Arizona-Utah border...15
15 Strial punctures medium sized, larger than those of intervals…16
15’ Strial punctures smaller, subequal to those of intervals…17
16 Anterolateral angles of pronotum projected, Sky Islands in southeast Arizona...E. dissimilis
16’ Anterolateral angles not projected, truncate, the Grand Canyon...E. leptoscelis
17 Size smaller, Sonoran Desert including the Colorado Desert...E. delicata (E. nevadensis)
17’ Size larger, northern Arizona and southern Utah, usually in forested areas...E. wheeleri
18 Fairly large tubercles present on elytra...19
18’ Elytra without large tubercules, or tubercules quite small...20
19 Range in southern California to Oregon...E. granosa
19’ Range near the Four Corners...E. leechi
20 Pronotum evenly tuberculate, Oregon...E. spoliata
20’ Pronotum punctate, widespread...21
21 Antennas robust, body generally more robust...E. hirsuta
21’ Antennas thin, body generally more slender...E. pilosa
22 Size small or sometimes medium, covered in long setae, restricted to coastal and sometimes inland California, and Baja California, or rarely in the direction of and in Reno, or the Sonoran Desert...23
22’ Size small or large, usually glabrous, if with conspicuous setae then not found in coastal California, nor in Baja California, or the Sonoran Desert, and if found near Reno, Nevada, then third antennomere not shortened...28
23 Males with dentate profemora and elytra caudate, Sonoran Desert...E. tribulus
23’ Males nor females ever caudate or attenuate, coastal California and Baja California...24
24 Third antennomere shortened, body smaller and more slender, pill shaped, setae yellow, coastal California and towards and into western Nevada...E. littoralis
24’ Third antennomere not shorted, body larger, California...25
25 Size 13mm or more...26
25’ Size 12mm or less...27
26 Setae usually orange, pronotum constricted at base, California and Baja California...E. osculans
26’ Setae black or grey, pronotum not constricted at base, Baja California...E. ursus
27 Setae black, pronotum constricted at base, California and Baja California...E. nigropilosa
27’ Setae yellow, pronotum not constricted at base, Baja California...E. subdeplanata
28 Elytra caudate at least in males and strongly muricate, near Tamaulipas if not caudate, size usually large...29
28’ Elytra caudate or not, but if caudate and muricately punctured then elytra only slightly muricate, elytra can be strongly muricate if not caudate and not near Tamaulipas, size large or small...31
29 Only males caudate with profemora dentate, Tamaulipas...E. fiski
29’ Both males and females caudate with mutic profemora, Four Corners and Sierra Nevada region...30
30 Elytra lacking long setae, Four Corners...E. caudifera
30’ Elytra always with long setae, Sierra Nevada region...E. longipilosa
31 Elytral sculpture very rough, elytra always attenuate but never caudate, pronotum wide, restricted to California, or slightly into Nevada and Oregon...32
31’ Elytral sculpture variable but if rough then elytra never attenuate and pronotum never wide, widespread...33
32 Luster shiny and elytral apex acutely attenuate, Pacific coast...E. marginata
32’ Luster dull and elytral apex blunt yet still attenuate, Sierra Nevada Mountains...E. scabricula (E. acutangula)
33 Body small and fusiform, punctures usually small and scattered, sometimes with minute setae, pronotum never sinuate before its base...34
33’ Shape rarely fusiform, if so, then punctures larger and in rows, pronotum sinuate before base, or size large...37
34 Elytra carinate and with minute setae…35
34’ Elytra without carina or minute setae...36
35 Line where pronotum and elytra meet nearly a perfect straight line, coastal Texas and Louisiana...E. veterator
35’ Line where pronotum and elytra meet rounded, making the elytra’s humeral angles appear projected, northern Texas to Canada...E. opaca
36 Body more fusiform, hind angles of pronotum wider than anterolateral angles...E. fusiformis
36’ Body less fusiform, hind angles and anterolateral angles about the same width...E. extricata (in part)
37 Size medium to slightly large, elytra usually inflated, wide, and large or sometimes flattened, especially in E. striolata, punctures either large, or appearing absent, although if elytra are not inflated, and attenuate, then punctures may be numerous and conspicuous, never sulcate...38
37’ Size variable but abdomen rarely inflated, if so, then punctures are small but conspicuous, sulcate or not...42
38 Elytral striae composed of large conspicuous punctures...E. goryi
38’ Elytral striae without large punctures...39
39 Elytra usually flattened, always attenuate, and heavily punctate, luster shiny, south and central Texas...E. striolata
39’ Elytra sometimes flattened, inconspicuous punctures present, although sometimes numerous and fairly noticeable, luster usually dull, Arizona, New Mexico, and Texas...40
40 Pronotum evenly arcuate and with anterolateral angles never projecting outwards, body grayish in color, covered in small punctures, Texas…E. spiculifera
40’ Pronotum parallel sided or evenly arcuate, anterolateral angles projecting outwards or not, color black, punctures inconspicuous, Arizona and New Mexico…41
41 Pronotum more or less parallel sided, anterolateral angles almost always projected outwards, prosternum forming large triangular wedge...E. subnitens
41’ Pronotum evenly arcuate, anterolateral angles only rarely projected outwards, prosternum not forming large triangular wedge...E. madrensis
42 Size almost always larger than 18mm, in rare cases reaching 14mm, but almost always with legs long, no matter the size, sometimes caudate, profemora often dentate, at least in males...43
42’ Size almost always smaller than 18mm, if larger then legs almost always short, unless body is fusiform, legs are often long if body in normal size range, never caudate, profemora only rarely dentate...72
43 Body elongate, elytra sometimes inflated, pronotum usually not all that distinct from elytra (subfusiform), if the pronotum and elytra are fairly distinct, then elytra smooth, punctures minute and simple, and prosternum projected downwards or profemora strongly sinuate, profemora never dentate and never with caudate elytra, punctures simple or muricate/scabrous, never sulcate...44
43’ Body fairly variable, sometimes elongate, often robust, pronotum usually quite distinct from elytra, almost always with dentate profemora, if missing then profemora slightly sinuate, elytra sometimes caudate, sulcate or not...49
44 Elytral punctures simple, anterolateral angles never very projected, body rarely flattened, but if so, then usually near the Sky Islands of southeast Arizona, size larger…46
44’ Elytral punctures not simple, anterolateral angles of pronotum projected, or body somewhat flattened and small, never in the Sky Islands, size smaller…45
45 Anterolateral angles strongly projected, elytral punctures with tiny inconspicuous setae arising from each one...E. inornata
45’ Anterolateral angles not projected or only slightly projected, elytral punctures glabrous…E. nigrina (continuation)
46 Elytra with red line running down the length of the elytra, this line never diffuses into black...E. ornatipennis
46’ Elytra entirely black, or if a red line is present along the elytra, it diffuses into black...47
47 Profemora strongly sinuate, pronotum widest slightly anterior of middle...E. hepburni
47’ Profemora simple, pronotum widest at middle...48
48 Prosternum smaller, only slightly projecting downwards, restricted to California...E. gigantea
48’ Prosternum larger prominently projecting downwards, widespread but not found in California...E. longicollis
49 Elytra with carina, if carina absent and the elytra either sulcate or with numerous irregular punctures, then anterolateral angles of the pronotum rounded off and the pronotum widest anterior of middle, pronotum concave or convex...50
49’ Elytra lacking carina, if elytra sulcate or with numerous irregular punctures, then anterolateral angles of the pronotum acute, or if angles not acute then pronotum widest at middle, pronotum convex...55
50 Pronotum concave...E. suturalis
50’ Pronotum convex...51
51 Elytra with carina...E. acuta
51’ Elytra without carina...52 (E. obscura)
52 Elytra with clear, prominent striae...53
52’ Elytra without clear, prominent striae...54
53 Elytra deeply sulcate, west of the Great Plains...E. obscura sulcipennis
53’ Elytra slightly sulcate, Great Plains...E. obscura obscura
54 Elytra with simple punctures, western Texas and southern New Mexico...E. obscura glabrisulca
54’ Elytra with muricate punctures, Four Corners...E. obscura dispersa
55 Elytra with large glabrous costae, intervals minutely hirsute, often coated with white debris, males are caudate, females are sometimes slightly caudate, legs long...E. mirabilis
55’ Elytra variable but without costae, caudate or not...56
56 Pronotum usually wide and strongly arcuate, humeral angles of elytra obsolete, never attenuate...E. grandicollis
56’ Pronotum variable, if wide, then elytral apex attenuate at least, humeral angles of elytra not obsolete...57
57 Elytra strongly sulcate, at least on elytral apex…58
57’ Elytra not strongly sulcate…59
58 Pronotum widest anterior to middle, anterolateral angles of pronotum acute, widespread…E. hispilabris
58’ Pronotum widest at middle, anterolateral angles of pronotum rounded off, Texas…E. composita
59 Restricted to California and Baja California, if not in California then tibia strongly curved, profemoral spine small and blunt, and in Nevada, northwest Arizona, or extreme southwest Utah...60
59’ Widespread, but not found in California, Baja California, Nevada, northwest Arizona, or southwest Utah...66
60 Pronotum two or more times wider than the head, often caudate...E. acuticauda
60’ Pronotum less then two times wider than the head, rarely caudate...61
61 Pronotum more or less parallel sided, widest at middle, Mohave and Colorado Deserts, Nevada and Arizona...62
61’ Pronotum not parallel sided, or if so then elytral apex strongly attenuate and with groove in between both elytra, if in the Mohave or Colorado Desert then pronotum widest anterior of middle…63
62 Tibia strongly curved, profemoral spines blunt, never caudate...E. subcylindrica
62’ Tibia not strongly curved, profemoral spines acute, males caudate...E. mexicana
63 Pronotum widest at middle...64
63’ Pronotum widest anterior of middle...65
64 Elytra glabrous, punctures minute, simple, luster usually shiny, elytral apex with groove in between the elytra, Baja California…E. discincta
64’ Elytra setose, punctures muricate, luster dull, elytral apex without groove in between the elytra, Channel Islands…E. adumbrata
65 Pronotum usually broad and strongly arcuate, widespread across California but absent near the U.S.-Mexico border…E. dentipes
65’ Pronotum more parallel sided, only slightly arcuate, luster more shiny, southern California…E. gracilis distans
66 Body usually flattened, elytral punctures simple, in rows, and more minute, never caudate, pronotum widest at middle, luster dull, Texas…E. knullorum
66’ Body usually not flattened, punctures variable, caudate or not, pronotum variable, if widest at middle then luster shiny, widespread…67
67 Elytral punctures irregular and strongly muricate, never caudate…E. sponsa
67’ Elytral punctures in rows and with simple punctures, or slightly asperate punctures, mainly at the elytral apex, elytra usually caudate, at least in males…68
68 Anterolateral angles of pronotum facing straight forward or only slightly projected outwards, hind angles not rounded, males caudate and with the metatibia sinuate, southern Arizona…E. eschscholtzii eschscholtzii
68’ If in southern Arizona then anterolateral angles of pronotum projecting outwards, or hind angles of pronotum appearing quite rounded…69
69 Hind angles of pronotum appearing rounded, size large, elytra often inflated and strongly convex, or fairly parallel sided…70 (E. spinipes)
69’ Hind angles of pronotum not rounded or only slightly rounded, size medium to large, elytra never inflated, often parallel sided…71
70 Elytra more or less parallel sided, west of the Pecos River…E. spinipes macrura
70’ Elytra ventricose, east of the Pecos River…E. spinipes ventricosa
71 Elytra caudate or strongly attenuate, El Paso region…E. tenuipes
71’ Elytra at most slightly attenuate…E. gracilis gracilis
72 Elytra with conspicuous setae…73
72’ Elytra glabrous, or only with short inconspicuous setae…77
73 Size less than 12mm, elytra shiny with complex sculpturing, and covered in golden hairs, areas with sandy soils generally around the Four Corners…E. barbata
73’ Size usually small but rarely less than 12mm, elytra usually dull, sometimes shiny, never with golden hairs, usually in deserts above the Sonoran Desert…74
74 Size larger, body more robust, northern Arizona and southern Utah…E. halli
74’ Size smaller, body more slender, generally around the Great Basin or the Great Sand Dunes of Colorado…75
75 Distributed only in the Great Sand Dunes National Park…E. hirtipennis
75’ Distributed generally around the Great Basin…76
76 Setae black, generally around the Great Basin…E. letcheri
76’ Setae brownish, in and around Utah…E. aspera
77 Elytra flat and widened, about the same width as and meeting with pronotum to form a continuous line…78
77’ Elytra usually not flat or widened, and never meeting with pronotum to form a line, elytra elongate or ovate…82
78 Body covered in minute setae, elytra with rows of tubercules, widespread…E. tricostata
78’ Body glabrous, elytra lacking rows of tubercules, Texas, Oklahoma, and New Mexico…79
79 Elytral sculpturing rough, striae vague to absent, luster generally dull…E. neomexicana
79’ Elytra with clear striae, luster shiny…80
80 Elytra with deeply impressed striae…E. pedinoides
80’ Elytra with slightly impressed striae…80 (E. wenzeli)
81 Punctures of elytra fine but easily noticeable…E. wenzeli speculicollis
81’ Punctures of elytra extremely fine and minute…E. wenzeli wenzeli
82 Both elytra with around three large smooth costae, legs often dark red, southern Utah…E. parowana
82’ Both elytra without at least three large costae…83
83 Size small, elytral luster generally shiny, elytra only with conspicuous irregular punctures, never with tubercules…E. extricata (continuation)
83’ Size small to medium, luster variable although generally dull if elytra with irregular punctures, said irregular punctures also smaller, elytra with punctures in rows or irregular, or with tubercules, papillae, or rugae…84
84 Elytra generally inflated, if not, then elytra more or less parallel sided, usually tuberculate, larger protibial spur extending almost to third tarsomere, Pacific Northwest…E. humeralis
84’ Elytra inflated or not, but if with tubercules then elytra not inflated and not parallel sided, if without tubercules then not found in the Pacific Northwest, or if in the Pacific Northwest then pronotum not clearly widest in front of middle…85
85 Elytra with papillae, pronotum wide, luster dull, California…E. papillosa
85’ Elytra without papillae, luster variable…86
86 Elytra with coarse transverse tuberculate rugae, luster shiny…E. corvina
86’ Elytra without transverse rugae, tuberculate or not, luster variable…87
87 Elytra irregularly tuberculate, tubercules larger, elytra noticeably rough, protibial spurs smaller…88
87’ Elytra not irregularly tuberculate, or if so then tubercules smaller, elytra less rough, and protibial spurs larger…89
88 Pronotum more or less arcuate and wider than long…E. granulata
88’ Pronotum more or less parallel sided and longer than wide…E. subtuberculata
89 Size small, pronotum evenly arcuate, punctures minute, luster dull, body generally fairly short, Arizona and New Mexico…E. arcuata
89’ Size variable, pronotum variable, but if evenly arcuate, then punctures more conspicuous, luster variable…90
90 Luster shiny, pronotum evenly arcuate, elytral punctures usually fairly large and in rows, outer protibial spur clearly larger than inner protibial spur, body usually ovate…E. debilis
90’ Luster variable but usually not too shiny, pronotum variable, punctures variable but rarely large, if protibial spurs disproportionate in size, then elytra with small punctures or tubercules and luster more dull…91
91 Longer protibial spur extending to almost to the third tarsomere, elytra tuberculate…92 (E. rileyi)
91’ Longer protibial spur extending only to apex of the first tarsomere, elytra never tuberculate…93
92 Elytra shiny with prominent striae, Four Corners…E. rileyi reducta
92’ Elytra dull without very prominent striae, widespread…E. rileyi rileyi
93 “…longer protibial spur parallel sided in basal half then tapering to a point at apex,” luster dull, punctures irregular if in California, if not then punctures in rows, pronotum generally around the same width as elytra…94
93’ “…longer protibial spur evenly tapering to a point from base to apex,” luster usually shiny, sometimes dull, punctures variable, if in California then luster usually dull and punctures with some semblance of rows, pronotum usually either wider or more narrow than elytra…96 (E. carbonaria)
94 Elytral punctures irregular, San Francisco region…E. quadricollis
94’ Elytral punctures organized in rows, generally around southeast Arizona…95 (E. anthracina)
95 Size larger, elytral punctures less fine, widespread…E. anthracina anthracina
95’ Size smaller, elytral punctures more fine, Chiricahua Mountains…E. anthracina lustrans
96 Strial punctures and those of intervals subequal in size, often diffusing, restricted to California…E. carbonaria omissa
96’ Punctures variable, but if in California then strial punctures larger than those of intervals and in rows…97
97 Body usually fairly fusiform, punctures conspicuous and in rows, east of the Pecos River and always below the southernmost edge of Arizona and New Mexico’s border…E. carbonaria soror
97’ Body only rarely fusiform, punctures variable, west of the Pecos River, or if not then above the southernmost edge of Arizona and New Mexico’s border…98
98 Punctures of elytra simple…100
98’ Punctures of elytra muricate…99
99 Strial punctures and those of intervals subequal in size, often diffusing, generally around northwest New Mexico…E. carbonaria knausii
99’ Strial punctures larger than those of intervals, striae in rows, often in ridges, widespread…E. carbonaria obsoleta
100 Body usually more fusiform, punctures more minute, size larger than 17mm, southern Arizona and New Mexico…E. carbonaria chihuahuaensis
100’ Body usually not fusiform, punctures more conspicuous, size 17mm or smaller, widespread…E. carbonaria carbonaria

Publicado el jueves, 08 de febrero de 2024 a las 07:05 AM por eleodesthermopolis eleodesthermopolis | 0 comentarios | Deja un comentario

lunes, 29 de enero de 2024

V3.4 A Key to the Eleodes Species of the United States and Canada OUTDATED

I have added some more help in the more difficult keys related to Melaneleodes, along with E. arcuata and E. debilis. I have also improved the Metablapylis section in case you're dealing with E. dissimilis. More info in my profile: https://www.inaturalist.org/people/6449260

The key on Google Drive (looks nicer and can be saved): https://drive.google.com/file/d/13gUFzNciTWTBsaBOQYSh-qPdg9NyBMCu/view?usp=drivesdk

1 Mesofemora and metafemora dentate or subdentate...2
1’ Mesofemora and metafemora never dentate...4
2 Pronotum widest anterior of middle, elytra with simple punctures, widespread...E. armata
2’ Pronotum widest at middle, elytral punctures simple or muricate, Baja California...3
3 Elytra usually appearing rough, metatibia more or less straight...E. moesta
3’ Elytra appearing smooth, metatibia curved or bent...E. femorata
4 Body small, pronotum generally cordate and constricted at base, elytral apex not attenuate or caudate, roughly sculptured, never sulcate, sometimes with short setae, usually distributed across beaches, forests, and mountains, rarely in arid regions...subgenus Blapylis
4’ Body small or large, pronotum usually not cordate although often slightly constricted at base, elytral apex attenuate, caudate or neither, roughly sculptured or not, sulcate or not, with setae or not, widespread across all environments...5
5 Legs long with unproportionately small body, robust or not, body often flattened if not robust, never sulcate, if legs shorter, then body elongate or robust and flattened, size smaller...6
5’ Legs long or short but with body fairly proportionate, only rarely flattened, sulcate or not, if legs fairly long, then not with conspicuous setae, or a small, elongate, and flattened body, size variable...22
6 Body generally slender or rarely robust, always flattened, pronotal and elytral sculpturing simple (except in E. nigrina), elytra usually more parallel sided, never with conspicuous setae...7
6’ Body, generally robust, never flattened, pronotal and elytral sculpturing complex, elytra usually more ventricose, usually with conspicuous setae...18
7 Elytral sculpturing muricate or scabrous, southern New Mexico to Canada, Sierra Nevada Mountains to Rocky Mountains...E. nigrina (in part)
7’ Elytral sculpturing simple, always below the northernmost edge of Colorado’s border...8
8 Range only in the Inyo White Mountains...9
8’ Range widespread but not in the Inyo White Mountains...10
9 Profemora with blunt tooth, anterolateral angles of pronotum blunt but projected, legs shorter...E. aalbui
9’ Profemora only sinuate, anterolateral angles of pronotum slightly projected, legs longer...E. microps
10 Range near western Texas and the Guadalupe Mountains...11
10’ Range in the Southwest United States excluding Texas and the Guadalupe Mountains...13
11 Probasitarsus with golden setae present, range in the Guadalupe Mountains...E. guadalupensis
11’ Probasitarsus lacking golden setae, only with orange spines, range near southwestern Texas...12
12 Mentum with finger-like process, pronotum lacking projected anterolateral angles, widest at middle, near Rio Grande river...E. labialis
12’ Mentum without finger-like process, pronotum with slightly projected anterolateral angles, Big Bend National Park...E. easterlai
13 Range in the Mohave and Colorado Deserts of California, pronotum usually evenly arcuate, strial punctures larger than those of intervals...E. californica
13’ Range in Arizona, Utah, and the Colorado Desert, pronotum rarely evenly arcuate, strial punctures variable, but if in the Colorado Desert then those of striae and intervals small and subequal in size...14
14 Range in the Sonoran Desert including the Colorado Desert or the most western portion of the Arizona-Utah border...15
14’ Range in the Sky Islands or the Colorado Plateau...16
15 Anterolateral angles strongly projected, caves by the most western point of the Arizona-Utah border…E. wynnei
15’ Anterolateral angles not strongly projected, the Sonoran Desert including the Colorado Desert…E. delicata (E. nevadensis)
16 Body elongate, northern Arizona and southern Utah...17
16’ Body not elongate, Sky Islands in southeast Arizona...E. dissimilis
17 Strial punctures larger than punctures of intervals, anterolateral edge of pronotum truncate, restricted to the Grand Canyon...E. leptoscelis
17’ Strial punctures subequal to punctures of intervals, anterolateral edge of pronotum not truncate, or only slightly truncate, northern Arizona and southern Utah...E. wheeleri
18 Large tubercles present on Elytra...19
18’ Elytra without large tubercules...20
19 Range in southern California to Oregon...E. granosa
19’ Range near the Four Corners...E. leechi
20 Pronotum evenly tuberculate, Oregon...E. spoliata
20’ Pronotum punctate, widespread...21
21 Antennas robust, body generally more robust...E. hirsuta
21’ Antennas thin, body generally more slender...E. pilosa
22 Size small or sometimes medium, covered in long setae, restricted to coastal and sometimes inland California, and Baja California, or rarely in the direction of and in Reno, or the Sonoran Desert...23
22’ Size small or large, usually glabrous, if with conspicuous setae then not found in coastal California, nor in Baja California, or the Sonoran Desert, and if found near Reno, Nevada, then third antennomere not shortened...28
23 Males with dentate profemora and elytra caudate, Sonoran Desert...E. tribulus
23’ Males nor females ever caudate or attenuate, coastal California and Baja California...24
24 Third antennomere shortened, body smaller and more slender, pill shaped, coastal California and towards and into Reno, Nevada...E. littoralis
24’ Third antennomere not shorted, body larger, California...25
25 Size 13mm or more...26
25’ Size 12mm or less...27
26 Setae orange, pronotum constricted at base, California and Baja California...E. osculans
26’ Setae black or grey, pronotum not constricted at base, Baja California...E. ursus
27 Setae black, pronotum constricted at base, California and Baja California...E. nigropilosa
27’ Setae yellow, pronotum not constricted at base, Baja California...E. subdeplanata
28 Elytra caudate at least in males and strongly muricate, near Tamaulipas if not caudate, size usually large...29
28’ Elytra caudate or not, but if caudate and muricately punctured then elytra only slightly muricate, elytra can be strongly muricate if not caudate and not near Tamaulipas, size large or small...31
29 Only males caudate with profemora dentate, Tamaulipas...E. fiski
29’ Both males and females caudate with mutic profemora, Four Corners and Sierra Nevada region...30
30 Elytra lacking long setae, Four Corners...E. caudifera
30’ Elytra always with long setae, Sierra Nevada region...E. longipilosa
31 Elytral sculpture very rough, elytra always attenuate but never caudate, pronotum wide, restricted to California, or slightly into Nevada and Oregon...32
31’ Elytral sculpture variable but if rough then elytra never attenuate and pronotum never wide, widespread...33
32 Luster shiny and elytral apex acutely attenuate, Pacific coast...E. marginata
32’ Luster dull and elytral apex blunt yet still attenuate, Sierra Nevada Mountains...E. scabricula (E. acutangula)
33 Body oval shaped, coated in minute setae, coastal Texas and Louisiana...E. veterator
33’ Body shape variable, if small and shaped slightly oval then not found in coastal Texas or Louisiana...34
34 Body small and fusiform, punctures usually small and scattered...35
34’ Body small or large, shape variable but usually not fusiform, if so, then punctures usually larger and in rows...37
35 Elytra carinate, covered in minute setae...E. opaca
35’ Elytra lacking carina, glabrous...36
36 Body more fusiform, central and southern Great Plains...E. fusiformis
36’ Body less fusiform, the Sierra Nevadas to the Rocky Mountains, and the Arizona Sky Islands to British Colombia, northern Great Plains...E. extricata (in part)
37 Size medium to large, elytra usually inflated, wide, and large or sometimes flattened, especially in E. striolata, punctures either large, or appearing absent, although if elytra are not inflated, and attenuate, then punctures may be numerous and conspicuous, never sulcate...38
37’ Size variable but abdomen rarely inflated, if so, then punctures are small but conspicuous, sulcate or not...42
38 Elytral striae composed of large conspicuous punctures...E. goryi
38’ Elytral striae without large punctures...39
39 Elytra usually flattened, always attenuate, and heavily punctate, luster shiny, south and central Texas...E. striolata
39’ Elytra sometimes flattened, inconspicuous punctures present, although sometimes numerous and fairly noticeable, luster usually dull, Arizona, New Mexico, and Texas...40
40 Pronotum evenly arcuate and with anterolateral angles never projecting outwards, body grayish in color, covered in small punctures, Texas…E. spiculifera
40’ Pronotum parallel sided or evenly arcuate, anterolateral angles projecting outwards or not, color black, punctures inconspicuous, Arizona and New Mexico…41
41 Pronotum more or less parallel sided, anterolateral angles almost always projected outwards, prosternum forming large triangular wedge...E. subnitens
41’ Pronotum evenly arcuate, anterolateral angles only rarely projected outwards, prosternum not forming large triangular wedge...E. madrensis
42 Size almost always larger than 17mm, in rare cases reaching 14mm, but almost always with legs long, no matter the size, sometimes caudate...43
42’ Size almost always smaller than 17mm, if larger then legs almost always short, unless body is fusiform, legs are often long if body in normal size range, never caudate...72
43 Body elongate, sometimes robust, pronotum usually not all that distinct from elytra (subfusiform), if the pronotum and elytra are fairly distinct, then elytra smooth, punctures minute and simple, and prosternum projected downwards or profemora strongly sinuate, profemora never dentate and never with caudate elytra, punctures simple or muricate/scabrous, never sulcate...44
43’ Body fairly variable, sometimes elongate, often robust, pronotum usually quite distinct from elytra, almost always with dentate profemora, if missing then profemora slightly sinuate, elytra sometimes caudate, sulcate or not...49
44 Elytral punctures simple, anterolateral angles never very projected, body rarely flattened, but if so, then usually near the Sky Islands of southeast Arizona…46
44’ Elytral punctures not simple, anterolateral angles of pronotum projected, or body somewhat flattened and small, never in the Sky Islands…45
45 Anterolateral angles strongly projected, elytral punctures with tiny inconspicuous setae arising from each one, size usually not small...E. inornata
45’ Anterolateral angles not projected or only slightly projected, elytral punctures glabrous, size smaller, sometimes quite small…E. nigrina (continuation)
46 Elytra with red line running down the length of the elytra, this line never diffuses into black...E. ornatipennis
46’ Elytra entirely black, or if a red line is present along the elytra, it diffuses into black...47
47 Profemora strongly sinuate, pronotum widest slightly anterior of middle...E. hepburni
47’ Profemora simple, pronotum widest at middle...48
48 Prosternum smaller, only slightly projecting downwards, restricted to California...E. gigantea
48’ Prosternum larger prominently projecting downwards, widespread but not found in California...E. longicollis
49 Elytra with carina, if carina absent and the elytra either sulcate or with numerous irregular punctures, then anterolateral angles of the pronotum rounded off and the pronotum widest anterior of middle, pronotum concave or convex...50
49’ Elytra lacking carina, if elytra sulcate or with numerous irregular punctures, then anterolateral angles of the pronotum acute, or if angles not acute then pronotum widest at middle, pronotum convex...55
50 Pronotum concave...E. suturalis
50’ Pronotum convex...51
51 Elytra with carina...E. acuta
51’ Elytra without carina...52 (E. obscura)
52 Elytra with clear, prominent striae...53
52’ Elytra without clear, prominent striae...54
53 Elytra deeply sulcate, west of the Great Plains...E. obscura sulcipennis
53’ Elytra slightly sulcate, Great Plains...E. obscura obscura
54 Elytra with simple punctures, western Texas and southern New Mexico...E. obscura glabrisulca
54’ Elytra with muricate punctures, Four Corners...E. obscura dispersa
55 Elytra with large glabrous costae, intervals minutely hirsute, often coated with white debris, males are caudate, females are sometimes slightly caudate, legs long...E. mirabilis
55’ Elytra variable but without costae, caudate or not...56
56 Pronotum usually wide, humeral angles of elytra obsolete, never attenuate...E. grandicollis
56’ Pronotum variable, if wide, then elytral apex attenuate at least, humeral angles of elytra not obsolete...57
57 Elytra strongly sulcate, at least on elytral apex…58
57’ Elytra not strongly sulcate…59
58 Pronotum widest anterior to middle, anterolateral angles of pronotum acute, widespread…E. hispilabris
58’ Pronotum widest at middle, anterolateral angles of pronotum rounded off, Texas…E. composita
59 Restricted to California and Baja California, if not in California then tibia strongly curved, profemoral spine small and blunt, and in Nevada, northwest Arizona, or extreme southwest Utah...60
59’ Widespread, but not found in California, Baja California, Nevada, northwest Arizona, or Utah...66
60 Pronotum two or more times wider than the head...E. acuticauda
60’ Pronotum less then two times wider than the head...61
61 Pronotum more or less parallel sided, widest at middle, Mohave and Colorado Deserts, Nevada and Arizona...62
61’ Pronotum not parallel sided, or if so then elytral apex with groove in between both elytra, if in the Mohave or Colorado Desert then pronotum widest anterior of middle…63
62 Tibia strongly curved, profemoral spines blunt, never caudate...E. subcylindrica
62’ Tibia not strongly curved, profemoral spines acute, males caudate...E. mexicana
63 Elytral punctures minute, pronotum nearly parallel sided, although sometimes slightly arcuate, elytral apex with groove in between both elytra, Baja California…E. discincta
63’ Elytral punctures not minute, or if so then pronotum wide…64
64 Pronotum usually broad and strongly arcuate, elytral striae less prominent, widespread across California but absent in the most southernmost parts of California…E. dentipes
64’ Pronotum more narrow, less prominently arcuate, elytral striae more prominent, southern California…65
65 Pronotum widest anterior of middle, luster more shiny, elytra with simple punctures, glabrous, striae very clear, mainland southern California…E. gracilis distans
65’ Pronotum widest at middle, luster more dull, elytra with muricate punctures and conspicuous setae, striae not clearly defined, Channel Islands…E. adumbrata
66 Body flattened, elytra and pronotum not too distinctly separated, or if so then elytra remain wide up until their humeral angles, Texas…E. knullorum
66’ Body usually not flat, elytra and pronotum clearly separate, if elytra remain wide up until their humeral angles, then elytra inflated, widespread…67
67 Elytra strongly muricate, never caudate…E. sponsa
67’ Elytra with simple punctures or only slightly asperate, mainly at the elytral apex, usually caudate, at least in males…68
68 Anterolateral angles of pronotum facing straight forward or only slightly projected outwards, hind angles not rounded, males caudate and with the metatibia sinuate, southern Arizona…E. eschscholtzii eschscholtzii
68’ If in southern Arizona then anterolateral angles of pronotum projecting outwards, or hind angles of pronotum appearing quite rounded…69
69 Hind angles of pronotum appearing rounded, size large, elytra often inflated and strongly convex, or fairly parallel sided…70 (E. spinipes)
69’ Hind angles of pronotum not rounded or only slightly rounded, size medium to large, elytra never inflated, often parallel sided…71
70 Elytra more or less parallel sided, west of the Pecos River…E. spinipes macrura
70’ Elytra ventricose, east of the Pecos River…E. spinipes ventricosa
71 Elytra caudate or strongly attenuate, El Paso region…E. tenuipes
71’ Elytra at most slightly attenuate…E. gracilis gracilis
72 Elytra with conspicuous setae…73
72’ Elytra glabrous, or only with short inconspicuous setae…77
73 Size less than 12mm, elytra shiny with complex sculpturing, and covered in golden hairs, areas with sandy soils generally around the Four Corners…E. barbata
73’ Size usually small but rarely less than 12mm, elytra usually dull, sometimes shiny, never with golden hairs, usually in deserts above the Sonoran Desert…74
74 Size larger, body more robust, northern Arizona and southern Utah…E. halli
74’ Size smaller, body more slender, generally around the Great Basin or the Great Sand Dunes of Colorado…75
75 Distributed only in the Great Sand Dunes National Park…E. hirtipennis
75’ Distributed generally around the Great Basin…76
76 Setae black, generally around the Great Basin…E. letcheri
76’ Setae brownish, in and around Utah…E. aspera
77 Elytra flat and widened, about the same width as pronotum, sides sometimes slightly rounded or with carina…78
77’ Pronotum usually not as wide as elytra, sides of elytra rounded evenly, generally elongate or ovate…82
78 Body covered with minute setae, usually appearing grey in color…E. tricostata
78’ Body glabrous…79
79 Elytral sculpturing rough, striae vague to absent, luster generally dull…E. neomexicana
79’ Elytra with clear striae, luster shiny…80
80 Elytra with deeply impressed striae…E. pedinoides
80’ Elytra with slightly impressed striae…80 (E. wenzeli)
81 Punctures of elytra fine but easily noticeable…E. wenzeli speculicollis
81’ Punctures of elytra extremely fine and minute…E. wenzeli wenzeli
82 Both elytra with at least three large smooth costae, legs often dark red, southern Utah…E. parowana
82’ Both elytra without at least three large costae…83
83 Size small, elytral luster generally shiny, elytra only with conspicuous irregular punctures, never with tubercules…E. extricata (continuation)
83’ Size small to medium, luster variable although generally dull if elytra with irregular punctures, said irregular punctures also smaller, elytra with punctures in rows or irregular, or with tubercules, papillae, or rugae…84
84 Elytra generally inflated, if not, then elytra more or less parallel sided, usually tuberculate, larger protibial spur extending almost to third tarsomere, Pacific Northwest…E. humeralis
84’ Elytra inflated or not, but if with tubercules then elytra not inflated and not parallel sided, if without tubercules then not found in the Pacific Northwest, or if in the Pacific Northwest then pronotum not clearly widest in front of middle…85
85 Elytra with papillae, pronotum wide, luster dull, California…E. papillosa
85’ Elytra without papillae, luster variable…86
86 Elytra with coarse transverse tuberculate rugae, luster shiny…E. corvina
86’ Elytra without transverse rugae, tuberculate or not, luster variable…87
87 Elytra irregularly tuberculate, tubercules larger, elytra noticeably rough, protibial spurs smaller…88
87’ Elytra not irregularly tuberculate, or if so then tubercules smaller, elytra less rough, and protibial spurs larger…89
88 Pronotum more or less arcuate and wider than long…E. granulata
88’ Pronotum more or less parallel sided and longer than wide…E. subtuberculata
89 Size small, pronotum evenly arcuate, punctures minute, luster dull, body generally fairly short, Arizona and New Mexico…E. arcuata
89’ Size variable, pronotum variable, but if evenly arcuate, then punctures more conspicuous, luster variable…90
90 Luster shiny, pronotum evenly arcuate, elytral punctures usually fairly large and in rows, outer protibial spur clearly larger than inner protibial spur, body usually ovate…E. debilis
90’ Luster variable but usually not too shiny, pronotum variable, punctures variable but rarely large, if protibial spurs disproportionate in size, then elytra with small punctures or tubercules and luster more dull…91
91 Longer protibial spur extending to almost to the third tarsomere, elytra tuberculate…92 (E. rileyi)
91’ Longer protibial spur extending only to apex of the first tarsomere, elytra never tuberculate…93
92 Elytra shiny with prominent striae, Four Corners…E. rileyi reducta
92’ Elytra dull without very prominent striae, widespread…E. rileyi rileyi
93 “…longer protibial spur parallel sided in basal half then tapering to a point at apex,” luster dull, punctures irregular if in California, if not then punctures in rows, pronotum generally around the same width as elytra…94
93’ “…longer protibial spur evenly tapering to a point from base to apex,” luster usually shiny, sometimes dull, punctures variable, if in California then luster usually dull and punctures with some semblance of rows, pronotum usually either wider or more narrow than elytra…96 (E. carbonaria)
94 Elytral punctures irregular, San Francisco region…E. quadricollis
94’ Elytral punctures organized in rows, generally around southeast Arizona…95 (E. anthracina)
95 Size larger, elytral punctures less fine, widespread…E. anthracina anthracina
95’ Size smaller, elytral punctures more fine, Chiricahua Mountains…E. anthracina lustrans
96 Strial punctures and those of intervals subequal in size, often diffusing, restricted to California…E. carbonaria omissa
96’ Punctures variable, but if in California then strial punctures larger than those of intervals and in rows…97
97 Body usually fairly fusiform, punctures conspicuous and in rows, east of the Pecos River and always below the southernmost edge of Arizona and New Mexico’s border…E. carbonaria soror
97’ Body only rarely fusiform, punctures variable, west of the Pecos River, or if not then above the southernmost edge of Arizona and New Mexico’s border…98
98 Punctures of elytra simple…100
98’ Punctures of elytra muricate…99
99 Strial punctures and those of intervals subequal in size, often diffusing, generally around northwest New Mexico…E. carbonaria knausii
99’ Strial punctures larger than those of intervals, striae in rows, often in ridges, widespread…E. carbonaria obsoleta
100 Body usually more fusiform, punctures more minute, size larger than 17mm, southern Arizona and New Mexico…E. carbonaria chihuahuaensis
100’ Body usually not fusiform, punctures more conspicuous, size 17mm or smaller, widespread…E. carbonaria carbonaria

Publicado el lunes, 29 de enero de 2024 a las 02:35 PM por eleodesthermopolis eleodesthermopolis | 0 comentarios | Deja un comentario

viernes, 26 de enero de 2024

V3.3 A Key to the Eleodes Species of the United States and Canada OUTDATED

I decided to add more details in the Cratidus section since I've seen observations of E. osculans with (seemingly) black hair, although I suspect it's the camera. I also updated couplet 43 since E. gigantea 's pronotum is variable, and E. hepburni could also be mistaken. More info in my profile: https://www.inaturalist.org/people/6449260

The key on Google Drive (looks nicer and can be saved): https://drive.google.com/file/d/1TLmQtIq5ydcJNgVLym8CAAg0LmhRb6VO/view?usp=drivesdk

1 Mesofemora and metafemora dentate or subdentate...2
1’ Mesofemora and metafemora never dentate...4
2 Pronotum widest anterior of middle, elytra with simple punctures, widespread...E. armata
2’ Pronotum widest at middle, elytral punctures simple or muricate, Baja California...3
3 Elytra usually appearing rough, metatibia more or less straight...E. moesta
3’ Elytra appearing smooth, metatibia curved or bent...E. femorata
4 Body small, pronotum generally cordate and constricted at base, elytral apex not attenuate or caudate, roughly
sculptured, never sulcate, sometimes with short setae, usually distributed across beaches, forests, and mountains,
rarely in arid regions...subgenus Blapylis
4’ Body small or large, pronotum usually not cordate although often slightly constricted at base, elytral apex
attenuate, caudate or neither, roughly sculptured or not, sulcate or not, with setae or not, widespread across all
environments...5
5 Legs long with unproportionately small body, robust or not, body often flattened if not robust, never sulcate, if legs
not abnormally long, then body elongate and flattened...6
5’ Legs long or short but with body fairly proportionate, only rarely flattened, sulcate or not, if legs fairly long, then
not with conspicuous setae, or a small, elongate, and flattened body...22
6 Body generally slender and flattened, pronotal and elytral sculpturing simple (except in E. nigrina), never with
conspicuous setae...7
6’ Body, generally robust, never flattened, pronotal and elytral sculpturing complex, usually with conspicuous
setae...18
7 Elytral sculpturing muricate or scabrous, southern New Mexico to Canada, Sierra Nevada Mountains to Rocky
Mountains...E. nigrina (in part)
7’ Elytral sculpturing simple, always below the northernmost edge of Colorado’s border...8
8 Range only in the Inyo White Mountains...9
8’ Range widespread but not in the Inyo White Mountains...10
9 Profemora with blunt tooth, anterolateral angles of pronotum blunt but projected, legs shorter...E. aalbui
9’ Profemora only sinuate, anterolateral angles of pronotum slightly projected, legs longer...E. microps
10 Range near western Texas and the Guadalupe Mountains...11
10’ Range in the Southwest United States excluding Texas and the Guadalupe Mountains...13
11 Probasitarsus with golden setae present, range in the Guadalupe Mountains...E. guadalupensis
11’ Probasitarsus lacking golden setae, only with orange spines, range near southwestern Texas...12
12 Mentum with finger-like process, pronotum lacking projected anterolateral angles, widest at middle, near Rio
Grande river...E. labialis
12’ Mentum without finger-like process, pronotum with slightly projected anterolateral angles, Big Bend National
Park...E. easterlai
13 Range in the Mohave and Colorado Deserts of California, pronotum usually evenly arcuate, strial punctures larger
than those of intervals...E. californica
13’ Range in Arizona, Utah, and the Colorado Desert, pronotum rarely evenly arcuate, strial punctures variable, but if
in the Colorado Desert then those of striae and intervals small and subequal in size...14
14 Range in the Sonoran Desert including the Colorado Desert or the most western portion of the Arizona-Utah
border...15
14’ Range in the Sky Islands or the Colorado Plateau...16
15 Anterolateral angles strongly projected, caves by the most western point of the Arizona-Utah border…E. wynnei
15’ Anterolateral angles not strongly projected, the Sonoran Desert including the Colorado Desert…E. delicata (E.
nevadensis)
16 Body elongate, northern Arizona and southern Utah...17
16’ Body not elongate, Sky Islands in southeast Arizona...E. dissimilis
17 Strial punctures larger than punctures of intervals, anterolateral edge of pronotum truncate, restricted to the
Grand Canyon...E. leptoscelis
17’ Strial punctures subequal to punctures of intervals, anterolateral edge of pronotum not truncate, or only slightly
truncate, northern Arizona and southern Utah...E. wheeleri
18 Large tubercles present on Elytra...19
18’ Elytra without large tubercules...20
19 Range in southern California to Oregon...E. granosa
19’ Range near the Four Corners...E. leechi
20 Pronotum evenly tuberculate, Oregon...E. spoliata
20’ Pronotum punctate, widespread...21
21 Antennas robust, body generally more robust...E. hirsuta
21’ Antennas thin, body generally more slender...E. pilosa
22 Size small or sometimes medium, covered in long setae, restricted to coastal and sometimes inland California, and
Baja California, or rarely in the direction of and in Reno, or the Sonoran Desert...23
22’ Size small or large, usually glabrous, if with conspicuous setae then not found in coastal California, nor in Baja
California, or the Sonoran Desert, and if found near Reno, Nevada, then third antennomere not shortened...28
23 Males with dentate profemora and elytra caudate, Sonoran Desert...E. tribulus
23’ Males nor females ever caudate or attenuate, coastal California and Baja California...24
24 Third antennomere shortened, body smaller and more slender, pill shaped, coastal California and towards and into
Reno, Nevada...E. littoralis
24’ Third antennomere not shorted, body larger, California...25
25 Size 13mm or more...26
25’ Size 12mm or less...27
26 Setae orange, pronotum constricted at base, California and Baja California...E. osculans
26’ Setae black or grey, pronotum not constricted at base, Baja California...E. ursus
27 Setae black, pronotum constricted at base, California and Baja California...E. nigropilosa
27’ Setae yellow, pronotum not constricted at base, Baja California...E. subdeplanata
28 Elytra caudate at least in males and strongly muricate, near Tamaulipas if not caudate, size usually large...29
28’ Elytra caudate or not, but if caudate and muricately punctured then elytra only slightly muricate, elytra can be
strongly muricate if not caudate and not near Tamaulipas, size large or small...31
29 Only males caudate with profemora dentate, Tamaulipas...E. fiski
29’ Both males and females caudate with mutic profemora, Four Corners and Sierra Nevada region...30
30 Elytra lacking long setae, Four Corners...E. caudifera
30’ Elytra always with long setae, Sierra Nevada region...E. longipilosa
31 Elytral sculpture very rough, elytra always attenuate but never caudate, pronotum wide, restricted to California, or
slightly into Nevada and Oregon...32
31’ Elytral sculpture variable but if rough then elytra never attenuate and pronotum never wide, widespread...33
32 Luster shiny and elytral apex acutely attenuate, Pacific coast...E. marginata
32’ Luster dull and elytral apex blunt yet still attenuate, Sierra Nevada Mountains...E. scabricula (E. acutangula)
33 Body oval shaped, coated in minute setae, coastal Texas and Louisiana...E. veterator
33’ Body shape variable, if small and shaped slightly oval then not found in coastal Texas or Louisiana...34
34 Body small and fusiform, punctures usually small and scattered...35
34’ Body small or large, shape variable but usually not fusiform, if so, then punctures usually larger and in rows...37
35 Elytra carinate, covered in minute setae...E. opaca
35’ Elytra lacking carina, glabrous...36
36 Body more fusiform, central and southern Great Plains...E. fusiformis
36’ Body less fusiform, the Sierra Nevadas to the Rocky Mountains, and the Arizona Sky Islands to British Colombia,
northern Great Plains...E. extricata (in part)
37 Size medium to large, elytra usually inflated, wide, and large or sometimes flattened, especially in E. striolata,
punctures either large, or appearing absent, although if elytra are not inflated, and attenuate, then punctures may
be numerous and conspicuous, never sulcate...38
37’ Size variable but abdomen rarely inflated, if so, then punctures are small but conspicuous, sulcate or not...42
38 Elytral striae composed of large conspicuous punctures...E. goryi
38’ Elytral striae without large punctures...39
39 Elytra usually flattened, always attenuate, and heavily punctate, luster shiny, south and central Texas...E. striolata
39’ Elytra sometimes flattened, inconspicuous punctures present, although sometimes numerous and fairly
noticeable, luster usually dull, Arizona, New Mexico, and Texas...40
40 Pronotum evenly arcuate and with anterolateral angles never projecting outwards, body grayish in color, covered
in small punctures, Texas…E. spiculifera
40’ Pronotum parallel sided or evenly arcuate, anterolateral angles projecting outwards or not, color black, punctures
inconspicuous, Arizona and New Mexico…41
41 Pronotum more or less parallel sided, anterolateral angles almost always projected outwards, prosternum forming
large triangular wedge...E. subnitens
41’ Pronotum evenly arcuate, anterolateral angles only rarely projected outwards, prosternum not forming large
triangular wedge...E. madrensis
42 Size almost always larger than 17mm, in rare cases reaching 14mm, but almost always with legs long, no matter the
size, sometimes caudate...43
42’ Size almost always smaller than 17mm, if larger then legs almost always short, unless body is fusiform, legs are
often long if body in normal size range, never caudate...72
43 Body elongate, sometimes robust, pronotum usually not all that distinct from elytra (subfusiform), if the pronotum
and elytra are fairly distinct, then elytra smooth, punctures minute and simple, and prosternum projected
downwards or prefemora strongly sinuate, profemora never dentate and never with caudate elytra, punctures
simple or muricate/scabrous, never sulcate...44
43’ Body fairly variable, sometimes elongate, often robust, pronotum usually quite distinct from elytra, almost always
with dentate profemora, if missing then profemora slightly sinuate, elytra sometimes caudate, sulcate or not...49
44 Elytral punctures simple, anterolateral angles never very projected, body rarely flattened, but if so, then usually
near the Sky Islands of southeast Arizona…46
44’ Elytral punctures not simple, anterolateral angles of pronotum projected, or body somewhat flattened and small,
never in the Sky Islands…45
45 Anterolateral angles strongly projected, elytral punctures with tiny inconspicuous setae arising from each one, size
usually not small...E. inornata
45’ Anterolateral angles not projected or only slightly projected, elytral punctures glabrous, size smaller, sometimes
quite small…E. nigrina (continuation)
46 Elytra with red line running down the length of the elytra, this line never diffuses into black...E. ornatipennis
46’ Elytra entirely black, or if a red line is present along the elytra, it diffuses into black...47
47 Profemora strongly sinuate, pronotum widest slightly anterior of middle...E. hepburni
47’ Profemora simple, pronotum widest at middle...48
48 Prosternum smaller, only slightly projecting downwards, restricted to California...E. gigantea
48’ Prosternum larger prominently projecting downwards, widespread but not found in California...E. longicollis
49 Elytra with carina, if carina absent and the elytra either sulcate or with numerous irregular punctures, then
anterolateral angles of the pronotum rounded off and the pronotum widest anterior of middle, pronotum concave
or convex...50
49’ Elytra lacking carina, if elytra sulcate or with numerous irregular punctures, then anterolateral angles of the
pronotum acute, or if angles not acute then pronotum widest at middle, pronotum convex...55
50 Pronotum concave...E. suturalis
50’ Pronotum convex...51
51 Elytra with carina...E. acuta
51’ Elytra without carina...52 (E. obscura)
52 Elytra with clear, prominent striae...53
52’ Elytra without clear, prominent striae...54
53 Elytra deeply sulcate, west of the Great Plains...E. obscura sulcipennis
53’ Elytra slightly sulcate, Great Plains...E. obscura obscura
54 Elytra with simple punctures, western Texas and southern New Mexico...E. obscura glabrisulca
54’ Elytra with muricate punctures, Four Corners...E. obscura dispersa
55 Elytra with large glabrous costae, intervals minutely hirsute, often coated with white debris, males are caudate,
females are sometimes slightly caudate, legs long...E. mirabilis
55’ Elytra variable but without costae, caudate or not...56
56 Pronotum usually wide, humeral angles of elytra obsolete, never attenuate...E. grandicollis
56’ Pronotum variable, if wide, then elytral apex attenuate at least, humeral angles of elytra not obsolete...57
57 Elytra strongly sulcate, at least on elytral apex…58
57’ Elytra not strongly sulcate…59
58 Pronotum widest anterior to middle, anterolateral angles of pronotum acute, widespread…E. hispilabris
58’ Pronotum widest at middle, anterolateral angles of pronotum rounded off, Texas…E. composita
59 Restricted to California and Baja California, if not in California then tibia strongly curved, profemoral spine small
and blunt, and in Nevada, northwest Arizona, or extreme southwest Utah...60
59’ Widespread, but not found in California, Baja California, Nevada, northwest Arizona, or Utah...66
60 Pronotum two or more times wider than the head...E. acuticauda
60’ Pronotum less then two times wider than the head...61
61 Pronotum more or less parallel sided, widest at middle, Mohave and Colorado Deserts, Nevada and Arizona...62
61’ Pronotum not parallel sided, or if so then elytral apex with groove in between both elytra, if in the Mohave or
Colorado Desert then pronotum widest anterior of middle…63
62 Tibia strongly curved, profemoral spines blunt, never caudate...E. subcylindrica
62’ Tibia not strongly curved, profemoral spines acute, males caudate...E. mexicana
63 Elytral punctures minute, pronotum nearly parallel sided, although sometimes slightly arcuate, elytral apex with
groove in between both elytra, Baja California…E. discincta
63’ Elytral punctures not minute, or if so then pronotum wide…64
64 Pronotum usually broad and strongly arcuate, elytral striae less prominent, widespread across California but
absent in the most southernmost parts of California…E. dentipes
64’ Pronotum more narrow, less prominently arcuate, elytral striae more prominent, southern California…65
65 Pronotum widest anterior of middle, luster more shiny, elytra with simple punctures, glabrous, striae very clear,
mainland southern California…E. gracilis distans
65’ Pronotum widest at middle, luster more dull, elytra with muricate punctures and conspicuous setae, striae not
clearly defined, Channel Islands…E. adumbrata
66 Body flattened, elytra and pronotum not too distinctly separated, or if so then elytra remain wide up until their
humeral angles, Texas…E. knullorum
66’ Body usually not flat, elytra and pronotum clearly separate, if elytra remain wide up until their humeral angles,
then elytra inflated, widespread…67
67 Elytra strongly muricate, never caudate…E. sponsa
67’ Elytra with simple punctures or only slightly asperate, mainly at the elytral apex, usually caudate, at least in
males…68
68 Anterolateral angles of pronotum facing straight forward or only slightly projected outwards, males caudate and
with the metatibia sinuate, southern Arizona…E. eschscholtzii eschscholtzii
68’ If in southern Arizona then anterolateral angles of pronotum projecting outwards, or hind angles of pronotum
appearing quite rounded…69
69 Hind angles of pronotum appearing rounded, size large, elytra often inflated and strongly convex, or fairly parallel
sided…70 (E. spinipes)
69’ Hind angles of pronotum not rounded or only slightly rounded, size medium to large, elytra never inflated, but
often parallel sided…71
70 Elytra more or less parallel sided, west of the Pecos River…E. spinipes macrura
70’ Elytra ventricose, east of the Pecos River…E. spinipes ventricosa
71 Elytra caudate or strongly attenuate, El Paso region…E. tenuipes
71’ Elytra at most slightly attenuate…E. gracilis gracilis
72 Elytra with conspicuous setae…73
72’ Elytra glabrous, or only with short inconspicuous setae…77
73 Size less than 12mm, elytra shiny with complex sculpturing, and covered in golden hairs, areas with sandy soils
generally around the Four Corners…E. barbata
73’ Size usually small but rarely less than 12mm, elytra usually dull, sometimes shiny, never with golden hairs, usually
in deserts above the Sonoran Desert…74
74 Size larger, body more robust, northern Arizona and southern Utah…E. halli
74’ Size smaller, body more slender, generally around the Great Basin or the Great Sand Dunes of Colorado…75
75 Distributed only in the Great Sand Dunes National Park…E. hirtipennis
75’ Distributed generally around the Great Basin…76
76 Setae black, generally around the Great Basin…E. letcheri
76’ Setae brownish, in and around Utah…E. aspera
77 Elytra flat and widened, about the same width as pronotum, sides sometimes slightly rounded or with carina…78
77’ Sides of elytra rounded evenly, generally elongate or circular…82
78 Body covered with minute setae, usually appearing grey in color…E. tricostata
78’ Body glabrous…79
79 Elytral sculpturing rough, striae vague to absent, luster generally dull…E. neomexicana
79’ Elytra with clear striae, luster shiny…80
80 Elytra with deeply impressed striae…E. pedinoides
80’ Elytra with slightly impressed striae…80 (E. wenzeli)
81 Punctures of elytra fine but easily noticeable…E. wenzeli speculicollis
81’ Punctures of elytra extremely fine and minute…E. wenzeli wenzeli
82 Both elytra with at least three large smooth costae, legs often dark red, southern Utah…E. parowana
82’ Both elytra without at least three large costae…83
83 Size small, elytral luster generally shiny, elytra only with conspicuous irregular punctures, never with tubercules…
E. extricata (continuation)
83’ Size small to medium, luster variable although generally dull if elytra with irregular punctures, said irregular
punctures also smaller, elytra with punctures in rows or irregular, or with tubercules, papillae, or rugae…84
84 Elytra generally inflated, if not, then elytra more or less parallel sided, usually tuberculate, larger protibial spur
extending almost to third tarsomere, Pacific Northwest…E. humeralis
84’ Elytra inflated or not, but if with tubercules then elytra not inflated and not parallel sided, if without tubercules
then not found in the Pacific Northwest, or if in the Pacific Northwest then pronotum not clearly widest in front
of middle…85
85 Elytra with papillae, pronotum wide, luster dull, California…E. papillosa
85’ Elytra without papillae, luster variable…86
86 Elytra with coarse transverse tuberculate rugae, luster shiny…E. corvina
86’ Elytra without coarse transverse rugae, tuberculate or not, luster variable…87
87 Elytra with large tubercules…88
87’ Elytra without large tubercules…89
88 Pronotum more or less arcuate and wider than long…E. granulata
88’ Pronotum more or less parallel sided and longer than wide…E. subtuberculata
89 Size small, pronotum evenly arcuate, punctures minute, body generally fairly short, Arizona and New Mexico…E.
arcuata
89’ Size variable, pronotum variable, but if evenly arcuate, then punctures more conspicuous…90
90 Luster shiny, elytral punctures usually fairly large and in rows, outer protibial spur clearly larger than inner
protibial spur…E. debilis
90’ Luster variable but usually not too shiny, punctures variable but rarely large, if protibial spurs disproportionate in
size, then elytra with small punctures or tubercules and or luster more dull…91
91 Longer protibial spur extending to almost to the third tarsomere…92 (E. rileyi)
91’ Longer protibial spur extending only to apex of the first tarsomere…93
92 Elytra shiny with prominent striae, Four Corners…E. rileyi reducta
92’ Elytra dull without very prominent striae, widespread…E. rileyi rileyi
93 “…longer protibial spur parallel sided in basal half then tapering to a point at apex”…94
93’ “…longer protibial spur evenly tapering to a point from base to apex”…96 (E. carbonaria)
94 Elytral punctures irregular, San Francisco region…E. quadricollis
94’ Elytral punctures organized in rows, generally around southeast Arizona…95 (E. anthracina)
95 Size larger, elytral punctures less fine, widespread…E. anthracina anthracina
95’ Size smaller, elytral punctures more fine, Chiricahua Mountains…E. anthracina lustrans
96 Strial punctures and those of intervals subequal in size, often irregularly patterned, restricted to California…E.
carbonaria omissa
96’ Punctures variable, but if in California then strial punctures larger than those of intervals and in rows…97
97 Body usually fairly fusiform, punctures conspicuous and in rows, east of the Pecos River and always below the
southernmost edge of Arizona and New Mexico’s border…E. carbonaria soror
97’ Body only rarely fusiform, punctures variable, west of the Pecos River, or if not then above the southernmost edge
of Arizona and New Mexico’s border…98
98 Punctures of elytra simple…100
98’ Punctures of elytra muricate…99
99 Strial punctures and those of intervals subequal in size, often irregularly patterned, generally around northwest
New Mexico…E. carbonaria knausii
99’ Strial punctures larger than those of intervals, striae in rows, often in ridges, widespread…E. carbonaria obsoleta
100 Body usually more fusiform, size larger than 17mm, southern Arizona and New Mexico…E. carbonaria
chihuahuaensis
100’ Body usually not fusiform, size 17mm or smaller, widespread…E. carbonaria carbonaria

Publicado el viernes, 26 de enero de 2024 a las 05:30 PM por eleodesthermopolis eleodesthermopolis | 0 comentarios | Deja un comentario

jueves, 25 de enero de 2024

V3.2 A Key to the Eleodes Species of the United States and Canada OUTDATED

I should definitely refrain from saying that an update won’t be the final one, but I believe this is nearly as good as this key can get. I have moved some things around, and added more details. I think it’s fitting that E. armata is the first species keyed out, with its reputation as one of the easiest species to ID, while E. carbonaria carbonaria is the last, as one of the most variable subspecies of E. carbonaria, often confused with E. debilis or other E. carbonaria subspecies, and various other vaguely similar species. More info in my profile: https://www.inaturalist.org/people/6449260

The key on Google Drive (looks nicer and can be saved): https://drive.google.com/file/d/1oWtrh5NuMuYoi9ruTAfPS-nqHMxTYrcN/view?usp=drivesdk

1 Mesofemora and metafemora dentate or subdentate...2
1’ Mesofemora and metafemora never dentate...4
2 Pronotum widest anterior of middle, elytra with simple punctures, widespread...E. armata
2’ Pronotum widest at middle, elytral punctures simple or muricate, Baja California...3
3 Elytra usually appearing rough, metatibia more or less straight...E. moesta
3’ Elytra appearing smooth, metatibia curved or bent...E. femorata
4 Body small, pronotum generally cordate and constricted at base, elytral apex not attenuate or caudate, roughly sculptured, never sulcate, sometimes with short setae, usually distributed across beaches, forests, and mountains, rarely in arid regions...subgenus Blapylis
4’ Body small or large, pronotum usually not cordate although often slightly constricted at base, elytral apex attenuate, caudate or neither, roughly sculptured or not, sulcate or not, with setae or not, widespread across all environments...5
5 Legs long with unproportionately small body, robust or not, body often flattened if not robust, never sulcate, if legs not abnormally long, then body elongate and flattened...6
5’ Legs long or short but with body fairly proportionate, only rarely flattened, sulcate or not, if legs fairly long, then not with conspicuous setae, or a small, elongate, and flattened body...22
6 Body generally slender and flattened, pronotal and elytral sculpturing simple (except in E. nigrina), never with conspicuous setae...7
6’ Body, generally robust, never flattened, pronotal and elytral sculpturing complex, usually with conspicuous setae...18
7 Elytral sculpturing muricate or scabrous, southern New Mexico to Canada, Sierra Nevada Mountains to Rocky Mountains...E. nigrina (in part)
7’ Elytral sculpturing simple, always below the northernmost edge of Colorado’s border...8
8 Range only in the Inyo White Mountains...9
8’ Range widespread but not in the Inyo White Mountains...10
9 Profemora with blunt tooth, anterolateral angles of pronotum blunt but projected, legs shorter...E. aalbui
9’ Profemora only sinuate, anterolateral angles of pronotum slightly projected, legs longer...E. microps
10 Range near western Texas and the Guadalupe Mountains...11
10’ Range in the Southwest United States excluding Texas and the Guadalupe Mountains...13
11 Probasitarsus with golden setae present, range in the Guadalupe Mountains...E. guadalupensis
11’ Probasitarsus lacking golden setae, only with orange spines, range near southwestern Texas...12
12 Mentum with finger-like process, pronotum lacking projected anterolateral angles, widest at middle, near Rio Grande river...E. labialis
12’ Mentum without finger-like process, pronotum with slightly projected anterolateral angles, Big Bend National Park...E. easterlai
13 Range in the Mohave and Colorado Deserts of California, pronotum usually evenly arcuate, strial punctures larger than those of intervals...E. californica
13’ Range in Arizona, Utah, and the Colorado Desert, pronotum rarely evenly arcuate, strial punctures variable, but if in the Colorado Desert then those of striae and intervals small and subequal in size...14
14 Range in the Sonoran Desert including the Colorado Desert or the most western portion of the Arizona-Utah border...15
14’ Range in the Sky Islands or the Colorado Plateau...16
15 Anterolateral angles strongly projected, caves by the most western point of the Arizona-Utah border…E. wynnei
15’ Anterolateral angles not strongly projected, the Sonoran Desert including the Colorado Desert…E. delicata (E. nevadensis)
16 Body elongate, northern Arizona and southern Utah...17
16’ Body not elongate, Sky Islands in southeast Arizona...E. dissimilis
17 Strial punctures larger than punctures of intervals, anterolateral edge of pronotum truncate, restricted to the Grand Canyon...E. leptoscelis
17’ Strial punctures subequal to punctures of intervals, anterolateral edge of pronotum not truncate, or only slightly truncate, northern Arizona and southern Utah...E. wheeleri
18 Large tubercles present on Elytra...19
18’ Elytra without large tubercules...20
19 Range in southern California to Oregon...E. granosa
19’ Range near the Four Corners...E. leechi
20 Pronotum evenly tuberculate, Oregon...E. spoliata
20’ Pronotum punctate, widespread...21
21 Antennas robust, body generally more robust...E. hirsuta
21’ Antennas thin, body generally more slender...E. pilosa
22 Size small or sometimes medium, covered in long setae, restricted to coastal and sometimes inland California, and Baja California, or rarely in the direction of and in Reno, or the Sonoran Desert...23
22’ Size small or large, usually glabrous, if with conspicuous setae then not found in coastal California, nor in Baja California, or the Sonoran Desert, and if found near Reno, Nevada, then third antennomere not shortened...28
23 Males with dentate profemora and elytra caudate, Sonoran Desert...E. tribulus
23’ Males nor females ever caudate or attenuate, coastal California and Baja California...24
24 Third antennomere shortened, body smaller and more slender, pill shaped, coastal California and towards and into Reno, Nevada...E. littoralis
24’ Third antennomere not shorted, body larger, California...25
25 Size 13mm or more...26
25’ Size 12mm or less...27
26 Setae orange, California and Baja California...E. osculans
26’ Setae black or grey, Baja California...E. ursus
27 Setae black, California and Baja California...E. nigropilosa
27’ Setae yellow, Baja California...E. subdeplanata
28 Elytra caudate at least in males and strongly muricate, near Tamaulipas if not caudate, size usually large...29
28’ Elytra caudate or not, but if caudate and muricately punctured then elytra only slightly muricate, elytra can be strongly muricate if not caudate and not near Tamaulipas, size large or small...31
29 Only males caudate with profemora dentate, Tamaulipas...E. fiski
29’ Both males and females caudate with mutic profemora, Four Corners and Sierra Nevada region...30
30 Elytra lacking long setae, Four Corners...E. caudifera
30’ Elytra always with long setae, Sierra Nevada region...E. longipilosa
31 Elytral sculpture very rough, elytra always attenuate but never caudate, pronotum wide, restricted to California, or slightly into Nevada and Oregon...32
31’ Elytral sculpture variable but if rough then elytra never attenuate and pronotum never wide, widespread...33
32 Luster shiny and elytral apex acutely attenuate, Pacific coast...E. marginata
32’ Luster dull and elytral apex blunt yet still attenuate, Sierra Nevada Mountains...E. scabricula (E. acutangula)
33 Body oval shaped, coated in minute setae, coastal Texas and Louisiana...E. veterator
33’ Body shape variable, if small and shaped slightly oval then not found in coastal Texas or Louisiana...34
34 Body small and fusiform, punctures usually small and scattered...35
34’ Body small or large, shape variable but usually not fusiform, if so, then punctures usually larger and in rows...37
35 Elytra carinate, covered in minute setae...E. opaca
35’ Elytra lacking carina, glabrous...36
36 Body more fusiform, central and southern Great Plains...E. fusiformis
36’ Body less fusiform, the Sierra Nevadas to the Rocky Mountains, and the Arizona Sky Islands to British Colombia, northern Great Plains...E. extricata (in part)
37 Size medium to large, elytra usually inflated, wide, and large or sometimes flattened, especially in E. striolata, punctures either large, or appearing absent, although if elytra are not inflated, and attenuate, then punctures may be numerous and conspicuous, never sulcate...38
37’ Size variable but abdomen rarely inflated, if so, then punctures are small but conspicuous, sulcate or not...42
38 Elytral striae composed of large conspicuous punctures...E. goryi
38’ Elytral striae without large punctures...39
39 Elytra usually flattened, always attenuate, and heavily punctate, luster shiny, south and central Texas...E. striolata
39’ Elytra sometimes flattened, inconspicuous punctures present, although sometimes numerous and fairly noticeable, luster usually dull, Arizona, New Mexico, and Texas...40
40 Pronotum evenly arcuate and with anterolateral angles never projecting outwards, body grayish in color, covered in small punctures, Texas…E. spiculifera
40’ Pronotum parallel sided or evenly arcuate, anterolateral angles projecting outwards or not, color black, punctures inconspicuous, Arizona and New Mexico…41
41 Pronotum more or less parallel sided, anterolateral angles almost always projected outwards, prosternum forming large triangular wedge...E. subnitens
41’ Pronotum evenly arcuate, anterolateral angles only rarely projected outwards, prosternum not forming large triangular wedge...E. madrensis
42 Size almost always larger than 17mm, in rare cases reaching 14mm, but almost always with legs long, no matter the size, sometimes caudate...43
42’ Size almost always smaller than 17mm, if larger then legs almost always short, unless body is fusiform, legs are often long if body in normal size range, never caudate...72
43 Body elongate, sometimes robust, pronotum not all that distinct from elytra, profemora never dentate and never with caudate elytra, punctures simple or muricate/scabrous, never sulcate...44
43’ Body fairly variable, sometimes elongate, often robust, pronotum usually quite distinct from elytra, males almost always with dentate profemora, elytra sometimes caudate, sulcate or not...49
44 Elytral punctures simple, anterolateral angles never very projected, body rarely flattened, but if so, then usually near the Sky Islands of southeast Arizona…46
44’ Elytral punctures not simple, anterolateral angles of pronotum projected, or body somewhat flattened and small, never in the Sky Islands…45
45 Anterolateral angles strongly projected, elytral punctures with tiny inconspicuous setae arising from each one, size usually not small...E. inornata
45’ Anterolateral angles not projected or only slightly projected, elytral punctures glabrous, size smaller, sometimes quite small…E. nigrina (continuation)
46 Elytra with red line running down the length of the elytra, this line never diffuses into black...E. ornatipennis
46’ Elytra entirely black, or if a red line is present along the elytra, it diffuses into black...47
47 Profemora strongly sinuate, pronotum widest slightly anterior of middle...E. hepburni
47’ Profemora simple, pronotum widest at middle...48
48 Prosternum smaller, only slightly projecting downwards, restricted to California...E. gigantea
48’ Prosternum larger prominently projecting downwards, widespread but not found in California...E. longicollis
49 Elytra with carina, if carina absent and the elytra either sulcate or with numerous irregular punctures, then anterolateral angles of the pronotum rounded off and the pronotum widest anterior of middle, pronotum concave or convex...50
49’ Elytra lacking carina, if elytra sulcate or with numerous irregular punctures, then anterolateral angles of the pronotum acute, or if angles not acute then pronotum widest at middle, pronotum convex...55
50 Pronotum concave...E. suturalis
50’ Pronotum convex...51
51 Elytra with carina...E. acuta
51’ Elytra without carina...52 (E. obscura)
52 Elytra with clear, prominent striae...53
52’ Elytra without clear, prominent striae...54
53 Elytra deeply sulcate, west of the Great Plains...E. obscura sulcipennis
53’ Elytra slightly sulcate, Great Plains...E. obscura obscura
54 Elytra with simple punctures, western Texas and southern New Mexico...E. obscura glabrisulca
54’ Elytra with muricate punctures, Four Corners...E. obscura dispersa
55 Elytra with large glabrous costae, intervals minutely hirsute, often coated with white debris, males are caudate, females are sometimes slightly caudate, legs long...E. mirabilis
55’ Elytra variable but without costae, caudate or not...56
56 Pronotum usually wide, humeral angles of elytra obsolete, never attenuate...E. grandicollis
56’ Pronotum variable, if wide, then elytral apex attenuate at least, humeral angles of elytra not obsolete...57
57 Elytra strongly sulcate, at least on elytral apex…58
57’ Elytra not strongly sulcate…59
58 Pronotum widest anterior to middle, anterolateral angles of pronotum acute, widespread…E. hispilabris
58’ Pronotum widest at middle, anterolateral angles of pronotum rounded off, Texas…E. composita
59 Restricted to California and Baja California, if not in California then tibia strongly curved, profemoral spine small and blunt, and in Nevada, northwest Arizona, or extreme southwest Utah...60
59’ Widespread, but not found in California, Baja California, Nevada, northwest Arizona, or Utah...66
60 Pronotum two or more times wider than the head...E. acuticauda
60’ Pronotum less then two times wider than the head...61
61 Pronotum more or less parallel sided, widest at middle, Mohave and Colorado Deserts, Nevada and Arizona...62
61’ Pronotum not parallel sided, or if so then elytral apex with groove in between both elytra, if in the Mohave or Colorado Desert then pronotum widest anterior of middle…63
62 Tibia strongly curved, profemoral spines blunt, never caudate...E. subcylindrica
62’ Tibia not strongly curved, profemoral spines acute, males caudate...E. mexicana
63 Elytral punctures minute, pronotum nearly parallel sided, although sometimes slightly arcuate, elytral apex with groove in between both elytra, Baja California…E. discincta
63’ Elytral punctures not minute, or if so then pronotum wide…64
64 Pronotum usually broad and strongly arcuate, elytral striae less prominent, widespread across California but absent in the most southernmost parts of California…E. dentipes
64’ Pronotum more narrow, less prominently arcuate, elytral striae more prominent, southern California…65
65 Pronotum widest anterior of middle, luster more shiny, elytra with simple punctures, glabrous, striae very clear, mainland southern California…E. gracilis distans
65’ Pronotum widest at middle, luster more dull, elytra with muricate punctures and conspicuous setae, striae not clearly defined, Channel Islands…E. adumbrata
66 Body flattened, elytra and pronotum not too distinctly separated, or if so then elytra remain wide up until their humeral angles, Texas…E. knullorum
66’ Body usually not flat, elytra and pronotum clearly separate, if elytra remain wide up until their humeral angles, then elytra inflated, widespread…67
67 Elytra strongly muricate, never caudate…E. sponsa
67’ Elytra with simple punctures or only slightly asperate, mainly at the elytral apex, usually caudate, at least in males…68
68 Anterolateral angles of pronotum facing straight forward or only slightly projected outwards, males caudate and with the metatibia sinuate, southern Arizona…E. eschscholtzii eschscholtzii
68’ If in southern Arizona then anterolateral angles of pronotum projecting outwards, or hind angles of pronotum appearing quite rounded…69
69 Hind angles of pronotum appearing rounded, size large, elytra often inflated and strongly convex, or fairly parallel sided…70 (E. spinipes)
69’ Hind angles of pronotum not rounded or only slightly rounded, size medium to large, elytra never inflated, but often parallel sided…71
70 Elytra more or less parallel sided, west of the Pecos River…E. spinipes macrura
70’ Elytra ventricose, east of the Pecos River…E. spinipes ventricosa
71 Elytra caudate or strongly attenuate, El Paso region…E. tenuipes
71’ Elytra at most slightly attenuate…E. gracilis gracilis
72 Elytra with conspicuous setae…73
72’ Elytra glabrous, or only with short inconspicuous setae…77
73 Size less than 12mm, elytra shiny with complex sculpturing, and covered in golden hairs, areas with sandy soils generally around the Four Corners…E. barbata
73’ Size usually small but rarely less than 12mm, elytra usually dull, sometimes shiny, never with golden hairs, usually in deserts above the Sonoran Desert…74
74 Size larger, body more robust, northern Arizona and southern Utah…E. halli
74’ Size smaller, body more slender, generally around the Great Basin or the Great Sand Dunes of Colorado…75
75 Distributed only in the Great Sand Dunes National Park…E. hirtipennis
75’ Distributed generally around the Great Basin…76
76 Setae black, generally around the Great Basin…E. letcheri
76’ Setae brownish, in and around Utah…E. aspera
77 Elytra flat and widened, about the same width as pronotum, sides sometimes slightly rounded or with carina…78
77’ Sides of elytra rounded evenly, generally elongate or circular…82
78 Body covered with minute setae, usually appearing grey in color…E. tricostata
78’ Body glabrous…79
79 Elytral sculpturing rough, striae vague to absent, luster generally dull…E. neomexicana
79’ Elytra with clear striae, luster shiny…80
80 Elytra with deeply impressed striae…E. pedinoides
80’ Elytra with slightly impressed striae…80 (E. wenzeli)
81 Punctures of elytra fine but easily noticeable…E. wenzeli speculicollis
81’ Punctures of elytra extremely fine and minute…E. wenzeli wenzeli
82 Both elytra with at least three large smooth costae, legs often dark red, southern Utah…E. parowana
82’ Both elytra without at least three large costae…83
83 Size small, elytral luster generally shiny, elytra only with conspicuous irregular punctures, never with tubercules…E. extricata (continuation)
83’ Size small to medium, luster variable although generally dull if elytra with irregular punctures, said irregular punctures also smaller, elytra with punctures in rows or irregular, or with tubercules, papillae, or rugae…84
84 Elytra generally inflated, if not, then elytra more or less parallel sided, usually tuberculate, larger protibial spur extending almost to third tarsomere, Pacific Northwest…E. humeralis
84’ Elytra inflated or not, but if with tubercules then elytra not inflated and not parallel sided, if without tubercules then not found in the Pacific Northwest, or if in the Pacific Northwest then pronotum not clearly widest in front of middle…85
85 Elytra with papillae, pronotum wide, luster dull, California…E. papillosa
85’ Elytra without papillae, luster variable…86
86 Elytra with coarse transverse tuberculate rugae, luster shiny…E. corvina
86’ Elytra without coarse transverse rugae, tuberculate or not, luster variable…87
87 Elytra with large tubercules…88
87’ Elytra without large tubercules…89
88 Pronotum more or less arcuate and wider than long…E. granulata
88’ Pronotum more or less parallel sided and longer than wide…E. subtuberculata
89 Size small, pronotum evenly arcuate, punctures minute, body generally fairly short, Arizona and New Mexico…E. arcuata
89’ Size variable, pronotum variable, but if evenly arcuate, then punctures more conspicuous…90
90 Luster shiny, elytral punctures usually fairly large and in rows, outer protibial spur clearly larger than inner protibial spur…E. debilis
90’ Luster variable but usually not too shiny, punctures variable but rarely large, if protibial spurs disproportionate in size, then elytra with small punctures or tubercules and or luster more dull…91
91 Longer protibial spur extending to almost to the third tarsomere…92 (E. rileyi)
91’ Longer protibial spur extending only to apex of the first tarsomere…93
92 Elytra shiny with prominent striae, Four Corners…E. rileyi reducta
92’ Elytra dull without very prominent striae, widespread…E. rileyi rileyi
93 “…longer protibial spur parallel sided in basal half then tapering to a point at apex”…94
93’ “…longer protibial spur evenly tapering to a point from base to apex”…96 (E. carbonaria)
94 Elytral punctures irregular, San Francisco region…E. quadricollis
94’ Elytral punctures organized in rows, generally around southeast Arizona…95 (E. anthracina)
95 Size larger, elytral punctures less fine, widespread…E. anthracina anthracina
95’ Size smaller, elytral punctures more fine, Chiricahua Mountains…E. anthracina lustrans
96 Strial punctures and those of intervals subequal in size, often irregularly patterned, restricted to California…E. carbonaria omissa
96’ Punctures variable, but if in California then strial punctures larger than those of intervals and in rows…97
97 Body usually fairly fusiform, punctures conspicuous and in rows, east of the Pecos River and always below the southernmost edge of Arizona and New Mexico’s border…E. carbonaria soror
97’ Body only rarely fusiform, punctures variable, west of the Pecos River, or if not then above the southernmost edge of Arizona and New Mexico’s border…98
98 Punctures of elytra simple…100
98’ Punctures of elytra muricate…99
99 Strial punctures and those of intervals subequal in size, often irregularly patterned, generally around northwest New Mexico…E. carbonaria knausii
99’ Strial punctures larger than those of intervals, striae in rows, often in ridges, widespread…E. carbonaria obsoleta
100 Body usually more fusiform, size larger than 17mm, southern Arizona and New Mexico…E. carbonaria chihuahuaensis
100’ Body usually not fusiform, size 17mm or smaller, widespread…E. carbonaria carbonaria

Publicado el jueves, 25 de enero de 2024 a las 06:19 AM por eleodesthermopolis eleodesthermopolis | 0 comentarios | Deja un comentario