Thallus: foliose, up to 5 cm in diam., usually +orbicular; lobes: discrete to more often somewhat irregularly flabellate, often confluent to imbricate, 0.5-1(-1.5) mm broad, +flat, prostrate; upper surface: gray to gray-brown, epruinose, rarely developing a patchy epinecral layer on lobe ends, without soredia or isidia but sometimes developing irregular lobules centrally; upper cortex: paraplectenchymatous; medulla: orange, sometimes white in first 0.5-1 mm of lobe ends; lower cortex: paraplectenchymatous; lower surface: black, often paler at the lobe ends, dull to weakly shiny; rhizines: simple, black; Apothecia: frequent and nearly always present, up to 2(-3) mm in diam., sessile; margin: usually crenate or becoming lobulate in older ones, often with a corona of rhizines; ascospores: ellipsoid, (15-)17.5-25.5 x 8-12.5 µm, of Physcia-type; Spot tests: all negative in medulla, medulla K+ purple; Secondary metabolites: skyrin, sometimes with a trace of zeorin.; Substrate and ecology: usually on rock, occasional on bark; World distribution: North America, Central America, Europe, Asia, and Africa; Sonoran distribution: mountains of Arizona, Chihuahua and Baja California Sur.; Notes: The distinctive red medulla of Phaeophyscia endococcinodes makes this species difficult to confuse with any other in the study area. This species is recognized as a variety of P. endococcina by Moberg (1985).